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SHELLF 


OF  THE 


South  Atl; 
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OFFICE    OF   T 

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OF    FISHERI 


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CONSERVATK 
BULLETIN 


UNITED  STATES  DEPARTMENT  OF  THE  INTERIOR 
Harold  L.  Ickes,  Secretary 

OFFICE  OF  THE  COORDINATOR  OF  FISHERIES 

Ira  N.  Gabrielson,  Deputy  Coordinator 


UNITED  STATES  GOVERNMENT  PRINTING  OFFICE  -  WASHINGTON  •  1944 
For  sale  by  the  Superintendent  of  Documents,  Washington  25,  D.  C.  —  Price  10  cents 


CONTENTS 


page 

Introduction 3 

The  fisheries 4 

The  fishing  grounds =. 6 

The  South  Atlantic  coast 6 

TheGulfcoast 7 

Boats,  men,  and  fishing  gear 8 

Marketing  the  catch 8 

Biographies  of  the  fish  and  shellfish  of  the  region 9 

Mullet 9 

Spanish  mackerel 12 

Kingfish L3 

Bluefish 14 

Red  snapper 1G 

Groupers 17 

Sea  trouts 19 

Redfish , 21 

Black  drum 22 

Shrimp 24 

The  oyster 20 

The  blue  crab 28 

Other  fish  and  shellfish 31 

Menhaden 31 

Shad 32 

Pompano 33 

Snook 34 

Grunts 35 

Sheepshead 35 

Blue  runner 37 

Crevalle_ 37 

Flounders 38 

Sharks 39 

Spiny  lobster 41 

Hard-shell  clam 41 

Bay  scallop 42 

Appendix 43 

The  nutritive  value  of  fish  and  shellfish ' 43 

General  guides  for  selecting  and  preparing  fish 43 

Bibliography 45 


Digitized  by  the  Internet  Archive 
in  2013 


http://archive.org/details/fishshellfishofsOOcars 


Fish  and  Shellfish 

of  the 

South  Atlantic  and  Gulf  Coasts 

By  Rachel  L.  Carson,  Aquatic  Biologist 
Office  of  the  Coordinator  of  Fisheries 


EVER  SINCE  the  first  settlements  were  made  in  the  region,  the  waters 
bordering  the  southern  Atlantic  coast  and  the  shores  of  the  Gulf  of 
Mexico  have  furnished  food.  Shrimp,  oysters,  crabs,  and  shorefish, 
abundant  in  the  warm  southern  bays  and  sounds,  insured  the  early 
colonists  against  starvation  and  later  formed  an  article  of  trade  with 
settlements  removed  from  the  coast,  upriver  and  in  the  foothills  of  the 
mountains.  Little  by  little  the  fisheries  of  the  region  developed,  and 
now  the  South  supplies  the  United  States  with  practically  all  of  its 
shrimp,  a  third  of  its  crabs,  a  fourth  of  its  oysters,  and  a  number  of 
choice  fishes  that  are  not  taken  anywhere  else  in  abundance,  like  the 
red  snapper,  the  Spanish  mackerel,  and  the  pompano. 

Although  mature  in  years,  the  fisheries  of  the  South  Atlantic  and 
Gulf  coasts  have  by  no  means  completed  their  development.  With  a 
coastline  more  than  2,500  miles  long  as  the  crow  flies,  the  region  produces 
only  a  little  more  than  half  a  billion  pounds  of  fishery  products — 14  per- 
cent of  the  total  yield  of  the  fisheries  of  the  United  States  and  Alaska. 
With  a  bewildering  variety  of  life  in  its  waters — more  kinds  of  fish  and 
shellfish  than  are  found  anywhere  else  in  the  country — the  region  has 
developed  only  five  major  fisheries:  the  menhaden,  shrimp,  mullet,  crab, 
and  oyster  industries.  Of  all  other  species  available,  none  are  caught 
in  quantities  exceeding  10,000,000  pounds. 

Part  of  this  slow  development  has  resulted  from  our  lack  of  knowledge, 
both  of  the  fishing  grounds  and  of  the  fish.  There  has  been  little  scien- 
tific exploration  of  the  region,  and  the  fishermen  have  confined  their 
operations  almost  entirely  to  the  waters  close  inshore.  We  do  not  know, 
with  few  exceptions,  the  areas  where  fish  may  congregate  offshore,  or 
whether  such  areas  exist.  We  know  very  little  of  the  life  histories  of 
some  of  the  most  important  species  of  the  region,  so  that  we  can  only 
guess  how  large  a  fishery  they  might  support.  Some  of  the  species  which 
we  know  to  be  abundant  cannot  now  be  fully  utilized  because  of  the 
limitations  of  the  present  methods  of  handling  the  catch. 

The  southern  fisheries  are  a  fertile  field  for  exploration  which  will 
reveal  whether  they  may  in  time  yield  an  even  larger  and  more  valuable 
seafood  harvest. 


CONSERVATION    BULLETIN    37 


THE   FISHERIES 


Cape  Hatteras  is  the  Mason  and  Dixon  Line  of  the  marine  world. 
North  of  it  lie  the  waters  inhabited  by  the  cold-water  fishes  of  the 
northern  ocean,  species  like  the  sea  herring,  the  common  mackerel,  cod, 
silver  hake,  and  pollock.  South  of  the  Cape  the  truly  southern  fauna 
begins.  Spanish  mackerel  and  kingfish,  mullet,  bluefish,  spotted  sea 
trout,  red  snapper,  groupers — these  are  a  few  of  the  many  species  that 
belong  to  the  warm  waters  of  the  South  and  are  caught  in  important 
quantities  nowhere  else.    To  be  sure,  many  of  them  make  occasional  or 


Table  1. — Fish  and  shellfish  landed  in  the  South  Atlantic  and  Gulf  states  in  19Jfi 
in  quantities  exceeding  100,000  pounds.1 


Pounds 


Value  to 
fishermen 


FISH 

Alewives  or  river  herring 

Bluefish 

Blue  runner 

Carp 

Catfish  and  bullhead 

Crappie 

Crevalle 

Croaker 

Drum: 

Black 

Red  or  redfish 

Flounders 

Groupers 

Harvestfish 

Hickory  shad 

Jewfish 

King  mackerel 

King  whiting 

Menhaden 

Mojarra 

Mullet 

Muttonfish 

Pinfish 

Pompano 

Sea  bass 

Sea  catfish 

Shad 

Sharks 

Sheepshead  (salt-water) 

Snappers: 

Mangrove 

Red 

Snook  or  sergeantfish 

Spanish  mackerel 

Spot 

Squeteagues  or  sea  trout: 

Gray 

Spotted 

White 

Striped  bass 

Sunfish 

Tenpounder 

White  perch 

Yellowtail 

SHELLFISH 

Crabs: 

Hard 

Soft  and  peelers 

Shrimp 

Spiny  lobster 

Clams,  hard 

Oysters,  market: 

Public,  spring 

Public,  fall 

Private,  spring 

Private,  fall 

Scallops,  bay 

Turtles,  soft-shell 


3 
1 

250 


37 


,115,900 

,438,400 
780 , 700 
438 , 000 

,486,200 
741,900 
238,700 

,421,400 

857,900 
,594,800 
,064,900 
, 184 . 000 
434 , 500 
371,500 
138,300 
,478,000 
,426,600 
,076,800 
394,900 
,078,100 
404,000 
130,100 
562 , 900 
420 , 800 
377 , 500 
,344,400 
,260,000 
628,900 

204.900 
,523,800 
470,000 
450,900 
,448,500 

630,500 
,311,900 
217,800 
539,900 
, 104 , 800 
227,100 
108,200 
59,800 


33,321,100 

543 , 700 

150,540,600 

463 , 500 

1,240,900 

12,742,100 
2,367,400 
3,391,900 
2,449,300 
162,200 
146,600 


$110,896 

178.166 

11,883 

19,662 

199,717 

23,857 

3 ,  385 

89,634 

37,945 

98,045 

83 , 555 

206,067 

12,735 

12,842 

4,381 

155,860 

41,721 

823 , 202 

6,688 

L, 378, 161 

23 , 098 

2,586 

137,321 

19,660 

7,056 

189,529 

25,200 

19,415 

9,359 
577,341 

24,519 
320,746 

85,989 

181,745 

512,207 

8,801 

59,389 

63,144 

5,052 

3,795 

5,338 


528,497 

83,311 

,893,360 

31,501 

112,638 

744,152 
185,372 
195,552 
137,674 
21,182 
2,204 


Source :  Fishery  Statistics  of  the  United  States,  1940.     Statistical  Digest  4,  Fish  and  Wildlife  Servic; 


FISH    AND    SHELLFISH  0 

even  regular  excursions  into  the  northern  area,  but  they  reach  their  full 
tide  of  abundance  only  along  southern  shores. 

.Because  it  lies  a  third  above  and  two  thirds  below  Cape  Hatteras, 
North  Carolina  has  a  fish  fauna  of  mixed  northern  and  southern  forms. 
Some  of  the  fish  taken  in  large  quantity  in  its  waters — such  as  the 
croaker,  spot,  alewives,  and  gray  sea  trout — also  are  found  in  great 
numbers  in  the  Chesapeake  Bay  and  along  the  Middle  Atlantic  coast. 
These  species  are  not  described  in  the  present  bulletin  because  they  are 
not  abundant  elsewhere  in  the  South  and  their  description  is  being 
reserved  for  a  future  publication  on  the  fishes  of  the  Middle  Atlantic 
coast  and  the  Chesapeake  Bay. 

In  1940,  a  year  fairly  representative  of  pre-war  conditions,  commercial 
fishermen  of  the  States  from  North  Carolina  to  Texas  brought  in  a  total 
of  573,533,000  pounds.  An  enormous  variety  of  edible  fish  and  shellfish 
were  represented  in  that  catch,  in  all  more  than  85  different  kinds.  But 
the  diversity  of  the  seafood  produced  by  the  southern  fisheries  is  more 
apparent  than  real.  Ninety  percent  of  the  entire  catch  consists  of  five 
species  of  aquatic  life,  while  the  remaining  10  percent  is  made  up  of 
75  species  caught  in  quantities  ranging  from  moderate  down  to  small 
or  insignificant.  The  menhaden  alone,  a  silvery  member  of  the  herring 
family  which  schools  in  enormous  numbers  at  the  surface,  comprises 
nearly  half  the  catch  of  all  species  for  the  entire  region.  Shrimp  con- 
tributes another  quarter.  Mullets,  crabs,  and  oysters  make  up  3  to 
6  percent  each.     (Detailed  statistics  of  the  catch  are  given  in  table  1.) 

As  might  be  expected,  the  eight  southern  coastal  States  differ  a  great 
deal  in  the  productivity  of  their  fisheries.  Florida,  with  its  long  coast- 
line, has  a  larger  catch  than  any  other  State  in  the  group,  contributing 
about  one-third  of  the  total  yield.  North  Carolina  is  a  close  second, 
with  30  percent  of  the  total,  and  Louisiana  is  third,  with  22  percent. 
Mississippi  furnishes  7  percent,  Texas  4,  and  Georgia,  Alabama,  and 
South  Carolina  1  to  2  percent  each.     (See  table  2  for  poundages.) 

Not  only  are  Florida's  fisheries  the  most  productive,  they  are  also 
the  most  varied,  drawing  on  67  species — 52  fish  and  15  shellfish.  This 
is  exclusive  of  a  number  of  fishes  like  the  tarpon,  bonefish,  and  others 
taken  only  by  sport  fishermen.  North  Carolina,  with  43  commercial 
species,  comes  next,  while  Mississippi,  drawing  on  only  17  species,  has 
the  least  diversified  fisheries  (table  2). 


Table  2. — Miscellaneous  information  on  the  fisheries  of  the  South  Atlantic  and 

Gulf  States.1 


State 

Num- 
ber of 
species 
in  com- 
mercial 

catch 

Total 

landings 

(in  pounds) 

Value  of 
catch  to 
fishermen 

Num- 
berof 
fisher- 
men 

Num- 
ber of 
ves- 
sels 2 

Largest 
fishery 
of  State 

Most 
valuable 
fishery  of 

State 

North  Carolina 

South  Carolina 

43 
21 
19 
67 
26 
17 
24 
22 

170,581,000 
9,445,900 

12,671,700 
187,492,200 

11,344,300 

38,002,300 
126,626,800 

19,369,200 

$1,864,164 
265 , 828 
380 , 630 

5,004,612 
561,911 
623,110 

4,951,493 
992,697 

7,113 
1,341 
1,178 
8,902 
1,134 
1,877 
4,858 
1,538 

184 
35 
77 

203 
41 

248 

288 
69 

Menhaden 

Crab 

Shrimp 

Menhaden 

Shrimp 

Menhaden 

Shrimp 

Shrimp 

Menhaden 

Oysters 

Shrimp 

Shrimp 
Shrimp 
Shrimp 

Mississippi 

Texas.. 

Total .. 

575 , 533 , 400 

14,644,895 

27,941 

1,145 

1  Based  on  statistics  for  1940. 

2  Exclusive  of  motor  boats,  row  boats,  and  other  small  craft. 


5  CONSERVATION    BULLETIN    3  7 

The  menhaden  industry  of  the  South  is  centered  in  North  Carolina, 
which  produces  slightly  more  than  half  the  total  yield  of  the  area. 
Louisiana  is  the  shellfish  capital  of  the  region,  with  the  southern  fisheries 
for  shrimp,  crabs,  and  oysters  all  most  highly  developed  in  that  State. 
Florida,  however,  is  the  headquarters  of  a  most  imposing  array  of  im- 
portant fisheries:  those  for  mullet,  red  snapper,  Spanish  mackerel,  king- 
fish,  bluefish,  sharks,  grouper,  and  redfish,  as  well  as  the  smaller  fisheries 
for  spiny  lobster,  pompano,  sheepshead,  blue  runner,  jacks,  and  clams. 

THE  FISHING  GROUNDS 

It  is  900  miles  from  Cape  Hatteras  to  the  Florida  Keys,  and  it  is 
1600  miles  from  the  Keys  around  the  Gulf  to  the  mouth  of  the  Rio 
Grande.  That  is  a  total  of  2,500  miles  of  coastline,  measured  directly 
and  without  following  all  the  small  irregularities,  the  numerous  islands, 
the  sounds  and  bays  and  marshes  that  make  the  South  Atlantic  and 
Gulf  coasts  the  shore  fisherman's  paradise.  Unlike  the  New  England 
fishermen,  who  take  most  of  their  catch  on  grounds  lying  far  offshore 
and  perhaps  hundreds  of  miles  from  the  home  port,  the  fishermen  of  the 
South  do  the  greater  part  of  their  fishing  in  a  narrow  band  of  water  only 
a  few  miles  wide  bordering  the  coast  on  its  ocean  side,  and  in  the  thou- 
sands of  square  miles  of  protected  bays  and  sounds  that  are  so  char- 
acteristic of  the  southern  coast. 

South  Atlantic  coast 

The  eastern  boundary  of  North  Carolina  is  a  chain  of  narrow  islands 
that  form  a  barrier  protecting  the  coast  from  the  pounding  of  the  Atlantic 
breakers.  Between  the  islands,  locally  called  "banks,"  and  the  coast  lie 
a  series  of  sounds — Currituck,  Albemarle,  Pamlico,  Core,  and  Bogue — 
some  3,000  square  miles  of  fishing  waters  connected  with  the  outside 
by  several  inlets.  Shad,  striped  bass,  and  river  herring  pass  through 
these  sounds  at  spawning  time,  and  sea  trout,  mullet,  croakers,  crabs, 
and  oysters  abound  in  them.  On  the  ocean  shore,  off  the  banks,  there 
is  good  fishing  for  other  species.  Famous  bluefish  grounds  lie  between 
Cape  Henry  and  Cape  Lookout.  Every  fall  the  spawning  runs  of  mullet 
pass  in  enormous  schools  along  the  outer  and  inner  shores  of  the  banks. 
Some  of  the  best  menhaden  grounds  are  in  the  waters  off  Cape  Lookout, 
and  sheepshead,  drum,  and  Spanish  mackerel  are  abundant  in  the  deep, 
protected  bight  west  of  the  Cape. 

Southward  along  the  coast  an  arc  of  almost  unbroken  beach  extends 
from  Cape  Fear  to  South  Carolina's  Winyah  Bay;  then  follows  a 
broken  coastline  of  marshes,  bays,  and  barrier  islands  through  the 
remainder  of  South  Carolina  and  Georgia,  nearly  to  the  St.  Johns  River. 
Then  the  broad,  sandy  beaches  of  eastern  Florida  sweep  in  a  long  curve 
to  the  Keys  and  the  beginning  of  the  Gulf  coast. 

This  much  of  the  South  Atlantic  coastline  can  be  seen  and  understood 
by  everyone.  We  know  much  less  about  what  lies  beneath  the  surface, 
the  undersea  hills  and  valleys  and  currents  that  determine  where  the 
fish  will  congregate  and  in  what  numbers.  Compared  with  offshore 
New  England — fished,  charted,  and  explored  since  the  days  of  John 
Cabot — the  South  Atlantic  and,  as  we  shall  see,  the  Gulf,  are  an  unknown 
marine  wilderness. 

We  do  know  that  the  continental  shelf,  the  gently  sloping  portion  of 
the  ocean  floor  that  extends  from  the  tide  lines  out  to  the  100-fathom 


FISH    AND    SHELLFISH  7 

depth,  is  only  about  30  miles  wide  off  Cape  Hatteras  and  averages  75 
to  100  miles  south  of  Hatteras  to  midway  down  the  Florida  coast,  when 
it  narrows  suddenly  to  a  mere  5  or  6  miles  off  Palm  Beach  and  Miami. 
Since  most  fish  are  confined  to  the  shallow  waters  over  the  continental 
shelf,  and  seldom  venture  out  over  true  oceanic  depths,  the  width  of  the 
shelf  has  a  direct  bearing  on  the  fisheries. 

Offshore  from  Hatteras  and  Lookout,  occasional  beds  of  coral  show 
the  beginning  of  a  truly  southern  fauna.  Off  the  coasts  of  South  Carolina 
and  Georgia  numerous  fishing  banks  are  scattered  over  the  continental 
shelf  between  the  10-fathom  and  100-fathom  line.  These  are  like  sub- 
marine hills  rising  above  a  deeper-lying  plain,  their  tops  covered  with 
sand,  coral  rock,  waving  sea  fans,  and  shells.  Sea  bass,  jacks,  flounders, 
snappers,  grunts,  and  sharks  are  some  of  the  species  fishermen  take  on 
the  southern  offshore  banks. 

Gulf  coast 

Off  the  southern  tip  of  Florida  a  long  chain  of  islands — the  Florida 
Keys — stretches  out  south  and  west  into  the  Gulf  like  giant  stepping 
stones.  Their  southern  border  swept  by  the  Gulf  Stream,  the  Keys  arc 
almost  inconceivably  rich  in  marine  life  of  all  kinds,  both  fishes  and 
the  smaller  invertebrate  life  they  feed  on.  Some  fish  are  more  abundant 
on  the  southern  side  of  the  reef  in  the  warmer  Gulf  Stream  water,  others 
on  the  north  side.  Not  infrequently  a  "norther"  drives  kingfish  and 
other  species  through  the  narrow  channels  between  the  reefs,  making 
good  fishing  while  the  schools  are  concentrated  in  a  restricted  area. 

Above  the  Keys,  the  west  coast  of  Florida  is  broken  into  a  maze  of 
waterways  about  the  "Ten  Thousand  Islands,"  then  swings  north  in 
a  long  arc,  interrupted  here  and  there  by  deeply  indenting  bays.  The 
continental  shelf  is  very  wide  off  this  coast — as  much  as  130  miles — and 
is  crossed  and  furrowed  by  ledges  and  gullies  where  fish  abound.  Its 
outer  margin  is  rocky  and  dotted  with  patches  of  coral. 

Westward,  bays,  sounds,  marshes,  and  chains  of  islands  mark  the 
coast  to  the  great  delta  of  the  Mississippi,  built  so  far  into  the  Gulf  that 
off  its  tip  the  continental  shelf  is  only  about  10  miles  wide.  Beyond  the 
Mississippi  are  more  coastal  marshes  for  mile  after  mile,  bordered  on 
the  seaward  side  by  barrier  beaches,  marked  by  vast  nearly  enclosed 
lagoons  and  bayous.  These  inside  waters  grow  fine  oysters,  and  they 
swarm  with  young  shrimp.  Redfish,  trout,  black  drum  and  other  fishes 
come  in  from  the  open  sea  as  young  fish  and  grow  and  mature  in  them. 

Of  the  open  Gulf  we  know  very  little,  for  it  is  a  region  that  has  been 
strangely  neglected  by  oceanographers,  and  with  the  single  exception 
of  the  combined  fishery  for  red  snappers  and  groupers,  the  Gulf  fisheries 
have  been  confined  to  a  narrow  band  of  coastal  waters.  The  area  of 
the  Gulf  is  about  700,000  square  miles.  Except  at  the  mouth  of  the 
Mississippi  and  again  at  Vera  Cruz,  the  continental  shelf  which  rims 
its  basin  is  wide — 100  to  130  miles.  The  deepest  part  of  the  Gulf's  floor 
— Sigsbee  Deep  off  the  coast  of  Yucatan — lies  more  than  two  miles 
below  the  surface,  and  much  of  the  remaining  area  is  from  one  to  two 
miles  deep.  The  Gulf  has  no  offshore  islands  except  the  Keys  of  Florida 
and  Yucatan,  and  the  island  of  Cuba.  Its  tides  have  a  range  of  only  two 
to  four  feet;  its  currents  sweep  in  a  vast  rotational  movement  about  its 
shores,  clockwise,  leaving  its  boundaries  by  way  of  the  Strait  of  Florida 
and  pouring  north  through  the  Atlantic  Ocean  as  the  river  of  warm 
blue  water  known  as  the  Gulf  Stream. 


g  CONSERVATION    BULLETIN    3  7 

BOATS,  MEN,  AND   FISHING  GEAR 

Before  the  war,  which  made  it  necessary  to  use  many  fishing  boats 
for  other  purposes,  the  fishing  fleet  of  the  South  Atlantic  and  Gulf 
coasts  was  the  largest  in  the  United  States  and  Alaska.  While  the 
number  of  large  vessels  was  slightly  below  the  number  of  similar  craft 
used  on  the  Pacific  coast  of  the  United  States — 1,145  compared  with 
1 ,563 — the  total  number  of  vessels,  motor  boats,  and  miscellaneous  small 
craft  fishing  out  of  southern  Atlantic  and  Gulf  ports  was  16,194.  The 
three  Pacific  Coast  States  had  the  next  largest  fleet,  with  9,105  craft, 
end  Alaska  the  third  largest  fleet  of  5,138. 

The  largest  vessels  operating  in  the  southern  area  are  the  menhaden 
purse  seiners,  which  cruise  about  in  the  larger  bays  or  in  the  open  ocean 
with  a  lookout  posted  in  the  crow's  nest  watching  for  schooling  men- 
haden. They  take  with  them  several  small  boats  from  which  fishermen 
carry  out  the  actual  operation  of  setting  the  seine-  around  the  fish.  The 
large  menhaden  seiners  probably  have  the  greatest  annual  production, 
per  boat,  of  any  vessels  operating  in  the  United  States  fisheries,  running 
from  7,000,000  to  10,000,000  pounds. 

Another  large  and  important  section  of  the  southern  fishing  fleet  is 
made  up  of  the  shrimp  trawlers,  which  bring  in  most  of  the  catch  of 
this  valuable  crustacean.  The  trawlers  drag  cone-shaped  nets  along 
the  bottom  of  the  sounds  or  ocean  where  the  shrimp  congregate  in  great- 
est numbers. 

Another  good  sized  fishing  vessel  seen  in  the  South  is  the  schooner 
type,  modeled  after  the  famous  Gloucester  schooners,  which  operates 
chiefly  out  of  Pensacola  and  visits  the  red  snapper  banks  off  the  coast 
of  Yucatan. 

A  variety  of  smaller  boats  is  used  in  operating  the  gill  nets,  haul 
seines,  and  dredges  which  are  important  in  many  southern  fisheries. 
Pound  nets,  common  in  North  Carolina,  but  not  farther  south,  are 
tended  usually  by  small  motor  boats.  Fyke  nets,  dip  nets,  and  cast 
nets  may  be  operated  from  the  shore. 

In  1940,  approximately  28,000  persons  were  directly  engaged  in  fishing 
in  the  States  from  North  Carolina  to  Texas.  Florida  reported  the  largest 
number  of  fishermen — 8,902.  North  Carolina  and  Louisiana  followed, 
with  7,113  and  4,858,  respectively.  Each  of  the  other  States  had  be- 
tween one  and  two  thousand  fishermen. 

MARKETING   THE  CATCH 

Although  the  South  Atlantic  and  Gulf  coasts  furnish  more  than  half 
a  billion  pounds  of  fish  and  shellfish  annually,  nearly  half  of  this  amount 
consists  of  menhaden,  practically  all  of  which  is  now  processed  into 
meal  and  oil  and  so  does  not  enter  the  ordinary  marketing  channels. 
Some  250,000,000  to  300,000,000  pounds  remain  for  distribution  as 
fresh,  frozen,  canned,  salted,  or  smoked  products. 

Most  of  the  fish  landed  at  southern  ports  enters  the  fresh-fish  markets. 
Despite  the  recent  introduction  here  and  there  of  the  modern  processing 
methods  of  preparing  fillets,  steaks,  and  tenderloins,  the  most  prevalent 
custom  is  to  ship  the  fish  in  the  round,  that  is,  without  preliminary 
dressing.  This  fact  makes  it  difficult  for  the  southern  States  to  compete 
with  New  England  as  a  fresh-fish  center,  for  in  that  area  much  of  the 
catch  is  routinely  processed  in  modern  plants  which  turn  out  a  con- 
veniently handled  product,  neatly  packaged  and  quick  frozen.  At 
present   the   necessary   shore   facilities — filleting   plants,    freezing   and 


FISH    AND    SHELLFISH  9 

storage  establishments,  and  the  like — are  extremely  limited  in  the 
South.  Likewise,  the  skilled  labor  necessary  to  operate  such  plants  is 
almost  unobtainable  under  war  conditions. 

The  piesent  methods  of  handling  the  catch  are  particularly  unsatis- 
factory as  applied  to  some  of  the  most  important  fisheries  of  the  region, 
such  as  the  mullet  and  Spanish  mackerel  fisheries.  Extremely  heavy 
catches  of  these  species  are  made  in  a  short  period,  so  that  in  practice 
it  has  often  proved  impossible  for  the  fresh  fish  markets  to  handle  all 
the  fish  that  are  caught.  The  problem  could  be  solved  if  means  were 
available  to  dress  the  fish  by  filleting  or  other  suitable  means,  quick 
freeze  it,  and  hold  it  for  distribution  in  accordance  with  market  demands. 

Canning  in  the  South  is  limited  almost  entirely  to  shrimp  and  oysters. 
It  has  been  the  practice  to  can  the  greater  part  of  the  shrimp,  and  this 
market  form  has  undoubtedly  been  most  familiar  throughout  the 
wide  area  in  which  shrimp  is  distributed,  but  there  is  now  a  strong 
trend  toward  freezing  more  shrimp  and  canning  less.  Practically  all 
the  oyster  canning  in  the  United  States  is  done  in  the  southern  States, 
and  Biloxi,  Miss.,  cans  more  oysters  than  any  other  city  in  the  world. 

Geographically,  the  products  of  the  southern  fisheries  (exclusive  of 
the  canned  pack)  are  distributed  chiefly  in  the  area  bounded  by  the 
Mississippi  River  on  the  west  and  the  Ohio  River  on  the  north.  Certain 
species,  however,  have  a  much  wider  distribution.  Probably  shrimp  and 
red  snapper  are  shipped  to  more  distant  cities  than  any  other  southern 
seafoods. 

From  the  South  Atlantic  coast  and  all  of  Florida  the  heaviest  ship- 
ments go  north  along  the  Atlantic  coast.  New  York  is  an  important 
market  for  this  area,  which  ships  as  far  north  as  Massachusetts,  Michi- 
gan, and  Illinois,  and  as  far  west  as  Texas  and  Kansas.  From  the 
Gulf  coast  many  shipments  go  north  by  truck  and  freight  up  the  Mis- 
sissippi Valley.  Shipments  are  also  made  to  the  Middle  Atlantic  area, 
the  New  England  States,  the  Rocky  Mountain  area,  and  the  Pacific 
coast. 

BIOGRAPHIES  OF  THE  FISH  AND  SHELLFISH  OF  THE  REGION 
Mullet  (Mugil  cephalus  and  M.  curema) 

The  mullet  is  the  most  important  food  fish  of  the  South.  It  supports 
the  most  valuable  fishery  in  the  States  of  Florida  and  South  Carolina, 
and  is  caught  in  larger  quantity  than  any  other  fish  with  the  exception 
of  menhaden  in  Florida,  Alabama,  and  Mississippi.  In  the  whole  area 
from  North  Carolina  to  Texas,  the  catch  of  mullet  is  exceeded  only 
by  menhaden,  which  is  used  primarily  for  the  manufacture  of  oil  and 
meal,  rather  than  as  food,  and  by  shrimp,  the  South 's  major  seafood 
product. 

Being  an  abundant  and  modestly  priced  fish,  mullet  is  one  of  the 
staple  foods  of  the  lower  income  groups  in  southern  coastal  cities. 
Despite  its  low  price,  it  is  one  of  the  better  food  fishes.  It  has  a  tender, 
firm-textured  flesh  which  contains  a  clear  yellow  oil  with  a  mild  and 
nutlike  flavor. 

Mullets  are  found  throughout  the  world  and  there  are  about  100 
different  species,  most  of  which  are  tropical.  The  most  abundant 
United  States  species,  the  striped  or  jumping  mullet  (Mugil  cephalus) 
itself  has  a  wide  distribution.  On  our  east  coast  it  occurs  all  the  way 
from  Cape  Cod  to  Brazil.     Occasionally  a  few  stray  mullet  appear 


10 


CONSERVATION    BULLETIN    3  7 


north  of  the  Cape,  in  the  Gulf  of  Maine,  but  such  an  occurrence  is  rare 
and  there  are  no  regular  fisheries  north  of  the  Carolinas.  On  the  Pacific 
coast  of  the  Americas  mullet  are  found  from  Monterey  to  Chile  and 
westwood  to  the  Hawaiian  Islands  and  Japan.  In  the  eastern  Atlantic 
they  are  common  on  the  southern  European  coast  and  in  the  Mediter- 
ranean. They  are  often  mentioned  in  the  writings  of  the  ancient 
Romans,  and  in  Egypt  they  have  been  cultivated  for  centuries  in  the 
overflow  deltas  of  the  Nile. 

The  white  mullet,  M.  cnrema,  is  taken  commercially  in  only  a  few 
localities,  although  it  occurs  all  the  way  from  Cape  Cod  to  Brazil.  It 
is  abundant  about  Key  West  and  the  other  Florida  Keys,  where  it 
makes  up  the  greater  part  of  the  mullet  catch.     It  is  so  similar  in 


Figure  1.— The  jumping  mullet  gets  its  name  from  its  habit  of  leaping  as  much  as  three  feet 
out  of  the  water,  making  it  easy  for  fishermen  to  spot  schools  of  this  fish. 


appearance  and  habits  to  the  striped  mullet  that  fishermen  do  not 
often  distinguish  between  them. 

In  their  relationships,  the  mullets  are  closest  to  the  silversides,  al- 
though much  larger  than  these  shimmering  little  minnows,  and  to  the 
barracudas.  The  striped  mullet  is  a  rather  stout  bodied  fish  with  large 
scales  and  averages  2  or  3  pounds  in  weight.  Probably  the  record 
size  is  10  pounds.  A  distinctive  mark  is  the  fact  that  it  has  two  well 
separated  dorsal  fins,  and  that  the  more  forwardly  placed  one  is  held 
stiffly  erect,  like  a  small  sail,  by  its  four  spines.  The  color  of  the  fish 
is  a  silvery  grey,  darker  on  the  back,  and  with  dusky  stripings  along  the 
sides.  Young  mullet  are  a  beautiful  silver  color,  like  a  freshly  minted 
dime. 

Mullet  live  in  coastal  waters  and  often  run  up  into  brackish  river 
mouths  to  feed.  In  South  America,  Dr.  Samuel  Hildebrand,  ichthy- 
ologist of  the  Fish  and  Wildlife  Service,  has  found  them  as  much  as 
one  hundred  miles  above  the  mouths  of  rivers,  sometimes  above  high 
waterfalls.  They  are  seldom  if  ever  found  in  fresh  water  in  the  United 
States. 

In  North  Carolina,  where  there  are  important  mullet  fisheries,  these 
fish  usually  spend  the  summer  in  the  bays  and  sounds,  feeding  on  the 
vegetation  of  the  bottom.  In  the  fall,  apparently  at  a  time  when  the 
spawn  is  maturing,  the  fish  begin  a  mass  exodus  from  these  inside 
waters,  passing  out  through  the  inlets  and  moving  southward  along  the 
coast  and  presumably  out  to  sea.  At  this  time  of  year  enormous  catches 
■ — sometimes  as  much  as  62,000  pounds  in  a  single  haul  of  the  net — are 
made  by  the  mullet  fishermen. 

Because  mullet  are  schooling  fish  and  especially  because  they  have 


FISH    AND    SHELLFISH  11 

a  remarkable  habit  of  jumping  as  much  as  three  feet  out  of  the  water, 
fishermen  can  easily  sight  an  approaching  school  and  set  their  nets 
about  it.  On  the  North  Carolina  coast  they  usually  wait  on  the  beach, 
boats  and  net  in  readiness,  until  the  lookout  on  top  of  a  nearby  dune 
gives  the  signal  that  the  fish  are  coming.  With  one  end  of  the  net 
held  on  shore,  the  seine  is  then  carried  out  in  a  large  semicircle  around 
the  fish  and  the  free  end  brought  in  to  the  beach.  The  men  then  jump 
out  of  the  boats  and  wade  out  into  the  surf  to  hold  down  the  lead  line 
in  the  lower  margin  of  the  net  with  their  feet,  otherwise  the  mullet  will 
escape  under  the  net.  At  best  a  number  of  them  will  leap  over  the 
cork  line,  sometimes  making  the  mistake  of  jumping  into  the  boats. 
Often  two  nets  are  set,  one  outside  the  other,  so  that  the  fish  that  escape 
the  inner  net  may  be  caught  in  the  outer.  The  net  is  drawn  in  as  rapidly 
as  possible  and  finally  the  central  part,  which  now  forms  a  sort  of 
bag  containing  the  fish,  is  hauled  up  on  the  beach,  all  the  mullet  flapping 
against  the  wet  sand  in  their  efforts  to  escape. 

In  Florida  mullet  are  caught  chiefly  at  night.  Although  unable  then 
to  see  the  fish  unless  the  night  is  bright,  fishermen  can  hear  a  school 
splashing  and  so  tell  when  to  set  the  net.  Gill  nets,  in  which  the  fish 
become  entangled  as  they  attempt  to  pass  through  the  meshes,  are  often 
used  in  Florida. 

Almost  three-fourths  of  the  entire  U.  S.  mullet  catch  of  35  to  40 
million  pounds  is  taken  in  Florida,  chiefly  on  the  west  coast.  Although 
far  below  Florida  in  production,  Alabama  ranks  second  in  the  mullet 
fishery,  followed  by  North  Carolina.  Mullet  are  present  in  abundance 
throughout  the  Gulf  of  Mexico,  but  they  have  been  almost  entirely 
neglected  in  Louisiana  and  Texas.  When  these  fisheries  are  developed, 
production  of  this  excellent  fish  can  be  greatly  increased  and  it  can 
be  supplied  in  larger  quantity  to  northern  cities. 

The  season  of  greatest  abundance  is  from  April  through  November, 
heaviest  runs  about  Beaufort,  N.  C,  occur  usually  in  September,  while 
the  peak  of  activity  in  the  Florida  fishery  comes  in  November.  In  that 
State  the  fishery  in  most  counties  is  closed  between  December  1  and 
January  10,  to  protect  the  fish  during  the  spawning  period. 

The  mullet  lends  itself  especially  well  to  cultivation,  and  has  been 
raised  in  ponds  for  many  centuries  in  various  parts  of  the  world.  The 
ancient  Romans  practiced  mullet  culture,  and  so  did  the  native  chief- 
tains of  the  Hawaiian  and  Philippine  Islands,  who  had  large  numbers 
of  fish  ponds  built  throughout  the  islands.  The  mullet  and  milkfish 
are  still  the  principal  species  raised  in  these  ponds. 

Some  recent  experiments  at  the  Beaufort,  N.  C,  laboratory  of  the  Fish 
and  Wildlife  Service  indicate  that  mullet  culture  in  shallow,  enclosed 
areas  along  the  South  Atlantic  coast  may  be  a  future  fish-cultural  de- 
velopment of  some  importance.  Dr.  Herbert  Prytherch,  in  charge  of  the 
laboratory,  says  that  mullet  which  he  placed  in  such  an  enclosure  as 
two-  or  three-inch  fish  grew  to  an  exceptional  size  in  two  years'  time. 
The  mullet  do  not  require  feeding,  provided  the  growth  of  vegetation 
is  stimulated  by  fertilization. 

At  present  mullet  usually  reach  the  market  in  the  round,  i.e.,  without 
being  dressed,  and  these  fish  may  be  either  fresh  or  frozen.  Some 
mullet  is  now  being  filleted  and  popular  demand  will  probably  lead  to 
more  of  them  being  prepared  in  this  convenient  form  after  the  war, 
when  labor  and  plant  facilities  permit.  Part  of  the  catch  is  salted,  this 
product  being  chiefly  for  export.    The  roe  also  is  eaten. 

Quite  a  range  of  sizes  is  available  for  the  purchaser — anything  from 
half-pound  fish,  which  are  best  suited  for  broiling,  to  large  five-pounders, 


12  CONSERVATION    BULLETIN    3  7 

which  should  be  baked.  The  Fish  and  Wildlife  Service  will  furnish,  on 
request,  a  leaflet  containing  seven  laboratory-tested  recipes  for  the 
preparation  of  mullet.1 

Spanish   Mackerel   (Scomberomorus   maculatus) 

The  Spanish  mackerel  is  predominantly  a  fish  of  warm  southern 
waters  but  despite  this  fact  its  distribution  is  astonishingly  wide.  It  may 
be  found  as  far  north  as  Maine  and  as  far  south  as  Brazil.  It  occurs  on 
the  far  side  of  the  Atlantic  in  waters  that  wash  the  African  coast  and  it 
lives  in  the  Pacific  from  San  Diego  to  the  Galapagos  Islands.  In  the 
United  States  only  a  few  thousand  pounds  are  taken  on  the  Pacific 
coast,  however,  and  although  commercial  catches  are  reported  on  the 
east  coast  all  the  way  from  New  Jersey  to  Texas,  the  real  center  of 
abundance  of  this  species  is  in  Florida. 


12,693 

Figure  2.— The  Spanish  mackerel   roves  the  warm  southern  oceans  in  search  of  its  prey,  the 

smaller  schooling  fishes. 

Only  a  little  more  than  a  hundred  years  ago,  the  Spanish  mackerel 
was  the  subject  of  one  of  the  shortest  biographies  ever  written.  "A  fine 
and  beautiful  fish;  comes  in  July,"  wrote  the  ichthyologist  Mitchell  in 
his  "Fishes  of  New  York,"  published  in  1815.  Even  in  the  late  1800's 
not  a  great  deal  could  be  added  to  this  one-sentence  description.  The 
Spanish  mackerel  was  known  only  as  an  excellent  food  fish  and  a  gamy 
fighter,  which  appeared  in  the  Middle  Atlantic  area  during  the  summer 
months,  retiring  in  the  fall  to  some  unknown  wintering  ground.  Where 
the  fish  goes  in  the  winter  is  no  longer  a  mystery,  for  most  of  Florida's 
6,000,000  pound  catch  is  made  between  November  and  March,  when 
the  Spanish  mackerel  is  absent  from  more  northern  waters.  It  is  now 
apparent  that  there  is  a  northward  migration  in  the  spring,  bringing  it 
into  the  North  Carolina  area  during  April  and  into  the  lower  Chesapeake 
Bay  by  May  or  June.  With  the  first  signs  of  fall  and  chilling  waters, 
however,  it  returns  to  the  more  comfortable  south. 

Unlike  the  groupers,  snappers,  and  drums,  which  lurk  in  coral  patches 
or  roam  over  deep-lying  ocean  bottoms,  the  Spanish  mackerel  live  almost 
entirely  at  the  surface.  Traveling,  they  move  in  schools,  and  are  easily 
seen  at  the  surface  as  they  pursue  their  food  or  clear  the  water  in 
apparently  playful  leaps.  According  to  Goode  (1887)  "the  surface  of 
the  Chesapeake  or  the  Gulf  of  Mexico  is  sometimes  broken  up  for  miles 
by  the  movements  of  large  schools  of  these  fishes,  while  the  air  is  en- 
livened by  the  screaming  flocks  of  terns,  which  follow  them,  to  gather  up 
the  fragments  of  their  feasts." 

1  Fishery   Leaflet  48,   Striped   mullet   and   their   preparation   for   the  table.      Mimeographed,   4   pp. 
Fish  and  Wildlife  Service,  Department  of  the  Interior,  Chicago  54,   111. 


FISH    AND    SHELLFISH  13 

Schools  of  menhaden  are  the  principal  source  of  food  for  the  predatory 
Spanish  mackerel,  which  also  feed  on  any  other  schooling  species,  such 
as  mullet,  silversides,  and  the  common  Boston  mackerel  where  the  ranges 
of  the  two  species  overlap. 

This  is  considered  an  even  more  beautiful  fish  than  the  famed  dolphin. 
It  is  a  deep  blue  with  iridescent  tones  of  gold  and  purple  on  the  back  and 
sides  and  silvery  below,  while  along  the  flanks  are  three  irregular  rows 
of  gold  spots,  "like  new  gold  dollars,"  as  an  early  writer  put  it. 

When  they  are  youngsters,  only  an  ichthyologist  could  distinguish  the 
Spanish  mackerel  from  its  close  relatives  the  kingfishes  or  ceros  (Scom- 
beromorus  cavalla  and  S.  regalis)  but  the  adults  develop  characteristics 
which  make  it  fairly  easy  to  tell  them  apart.  The  Spanish  mackerel  is 
a  smaller  fish  than  either  of  the  others,  averaging  only  about  2  pounds, 
and  attaining  a  maximum  weight  of  20  to  25  pounds.  S.  regalis,  which 
is  abundant  in  this  country  only  about  the  Florida  Keys,  averages  5 
pounds  and  may  reach  35 ;  S.  cavalla  averages  7  and  reaches  a  maximum 
of  50  to  75.  Apart  from  size  differences,  if  a  fish  of  this  group  has  yellow 
spots  on  its  flanks  and  a  gradually  curving  lateral  line,  it  is  a  Spanish 
mackerel;  if  it  has  spots  and  one  or  two  continuous  black  lines  along  its 
sides  it  is  S.  regalis;  if  it  has  no  (or  at  most  a  scattered  few)  spots  and  a 
lateral  line  that  drops  abruptly  opposite  the  second  dorsal  fin,  it  is 
S.  cavalla. 

The  Spanish  mackerel  spawns  while  on  its  summer  migration,  at  least 
from  the  vicinity  of  the  Carolinas  to  Long  Island,  but  we  do  not  know 
whether  spawning  may  also  take  place  north  and  south  of  these  limits. 
Hildebrand  and  Cable  (1940)  found  larvae  only  a  few  days  old  in  the 
open  ocean  off  Beaufort,  N.  C,  from  late  June  until  early  September,  but 
none  in  the  inshore  waters.  The  lower  Chesapeake  Bay,  however,  is  a 
spawning  ground  for  Spanish  mackerel,  although  the  fish  are  not  as 
numerous  in  the  Bay  as  they  formerly  were. 

In  Florida,  where  almost  the  entire  commercial  catch  is  made,  fisher- 
men take  the  Spanish  mackerel  in  gill  nets  and  purse  seines.  Most  of  the 
gill  netting  is  done  at  night,  because  the  fish  are  unable  to  see  the  net  in 
the  darkness  and  are  more  likely  to  gill  themselves.  The  boats  carry 
powerful  searchlights  to  aid  the  crews  in  locating  the  fish.  When  the 
mackerel  are  sighted,  the  fishermen  set  their  nets  around  the  school  as 
quickly  as  possible,  marking  its  position  by  lanterns  mounted  on  cork 
floats.  The  next  step  is  to  frighten  the  fish  as  much  as  possible,  causing 
them  to  plunge  against  the  net  and  become  entangled  by  their  gill  covers 
in  its  meshes.  This  is  done  by  splashing  in  the  water  with  oars,  or  by 
rowing  directly  into  the  space  enclosed  by  the  nets.  Although  there  is 
nothing  to  prevent  the  fish  from  swimming  deep  and  escaping  under  the 
net,  their  habits  are  such  that  they  ordinarily  remain  at  the  surface, 
either  gilling  themselves  or  escaping  over  the  cork  line. 

As  long  as  Spanish  mackerel  have  been  caught,  they  have  been  con- 
sidered one  of  the  choicest  species  of  the  east  coast,  ranking  close  to  the 
pompano  in  popular  esteem.  They  are  usually  marketed  whole,  and  are 
best  broiled  or  baked.  Shipments  are  made  throughout  eastern  United 
States  with  New  York  one  of  the  most  important  markets. 

Kingfish   (Scomberomorus   cavalla) 

The  kingfish,  giant  relative  of  the  prized  Spanish  mackerel,  takes  top 
rank  among  the  game  fishes  of  America  and  is  only  a  little  less  valued 
than  its  relative  as  a  food  fish.     It  occurs  in  great  numbers  at  certain 


14  CONSERVATION    BULLETIN    3  7 

seasons  off  the  coast  of  Florida  and  rarely  is  seen  as  far  north  as  Cape 
Qod.     Southward,  the  limit  of  its  range  is  Brazil. 

The  only  important  production  center  for  the  kingfish  in  the  United 
States  is  Florida,  where  three  to  three  and  a  half  million  pounds  are 
taken.  A  very  small  catch  is  made  in  Texas.  In  Florida  the  related 
species,  S.  regalis,  taken  chiefly  about  the  Keys,  is  also  called  the  kingfish 
and  fishermen  seldom  distinguish  between  them. 

The  season  for  kingfish  in  Florida  is  about  the  same  as  that  for  the 
Spanish  mackerel,  or  from  November  to  March.  It  is  taken  along  both 
the  Atlantic  and  Gulf  coasts,  chiefly  by  trolling,  and  is  found  in  large 
schools,  ranging  the  waters  of  the  open  sea  in  search  of  menhaden  and 
other  small  fish.    It  is  noted  for  its  remarkable  leaps,  often  clearing  the 


Figure  3. — The  kingfish  is  a  favorite  of  Southern  sport  fishermen  and  also  a  choice  food  fish. 

water  by  10  feet  or  more.  Fishermen  seeking  Spanish  mackerel  often 
take  the  kingfish  incidentally,  because  of  the  similar  habits  of  the  two 
species,  but  it  is  so  strong  and  has  such  sharp  teeth  that  it  does  consid- 
erable damage  to  the  gear. 

Kingfish  taken  by  trolling  usually  run  from  15  to  30  pounds  in  weight, 
but  the  maximum  size  for  the  species  is  about  75  pounds.  Its  size, 
strength,  speed,  and  great  fighting  powers  make  it  one  of  the  most 
important  game  fishes  on  the  coasts  of  Florida  and  North  Carolina. 

Like  the  Spanish  mackerel,  it  is  a  fish  of  fine  flavor,  with  few  bones. 
The  larger  fish  are  sold  in  steaks,  the  smaller  ones  whole.  Most  of  the 
catch  is  marketed  along  the  eastern  seaboard,  Richmond  and  New  York 
being  especially  important  markets  for  this  species. 

Bluefish  (Pomatomus  saltatrix) 

The  bluefish  is  another  warm-water  species  which  migrates  up  and 
down  the  Atlantic  coast  from  Florida  to  Massachusetts  following  the 
schools  of  menhaden  and  other  small  fish  and  avoiding  chilly  climates. 
In  midwinter  bluefish  are  to  be  found  in  southern  Florida,  where  the 
bulk  of  the  commercial  catch  is  made.  Like  migrant  birds,  they  turn 
north  with  the  spring,  and  by  March  and  April  large  schools  are  passing 
the  coasts  of  the  Carolinas.  They  reach  Long  Island  in  late  April  or 
early  May,  Massachusetts  by  late  May.  In  some  years,  probably  when 
they  are  especially  abundant,  they  pass  Cape  Cod  and  appear  during 
the  summer  months  in  Massachusetts  Bay  and  as  far  north  as  Maine, 
but  these  waters  are  beyond  the  range  of  their  normal  summer 
wanderings. 

The  total  United  States  catch  of  bluefish  was  only  2,629,000  pounds  in 
1940,  the  most  recent  year  for  which  we  have  a  complete  record.  This 
was  the  low  point  of  a  catch  that  has  been  steadily  dropping  since  1936, 
when  nearly  10,000,000  pounds  were  taken.  The  entire  history  of  the 
fishery  for  bluefish  is  one  of  rapid  changes  in  abundance.    Probably  the 


FISH    AND    SHELLFISH 


15 


largest  catches — some  20,000,000  pounds — were  made  about  1900,  but 
during  several  periods  the  yield  has  fallen  as  low  as  the  present  level. 
We  know  too  little  about  the  natural  history  of  the  bluefish  to  be  able 
to  give  a  satisfactory  explanation  of  these  changes  in  its  abundance. 
In  lean  bluefish  years  it  is  the  custom  for  sport  fishermen  to  say  that 
scarcity  of  menhaden,  probably  the  favorite  food  of  the  bluefish,  is 
responsible.  This  supposition  has  never  been  proved  and  the  records  do 
not  show  that  the  changes  in  abundance  of  the  two  species  coincide. 
A  more  probable  explanation  is  that  the  bluefish,  like  many  other  marine 
species,  is  subject  to  extreme  hazards  during  the  first  days  and  weeks 
of  life  from  unfavorable  winds,  currents,  and  water  temperatures,  as 
well  as  from  the  attacks  of  natural  enemies.    In  the  case  of  the  Atlantic 


w 


Figure  4.— The  bluefish,  a  strong,   swift  fish   of  the  open  sea,   may  run   up  to  three   feet   in 
length  and  is  a  deep  bluish  green  above  with  silver  below. 

mackerel  it  has  been  proved  that  if,  during  the  brief  spawning  period, 
the  water  is  unseasonably  cold  or  the  set  of  winds  and  currents  is  such 
as  to  carry  the  young  fish  out  of  their  normal  nursery  areas,  practically 
the  entire  brood  may  perish.  If  this  happens  two  or  three  years  in 
succession,  there  is  an  acute  shortage  of  mackerel.  Perhaps  something 
very  like  this  happens  in  the  life  of  the  bluefish,  but  we  shall  have  to 
wait  until  we  know  more  of  its  life  history  before  we  can  explain  the 
ups  and  downs  of  abundance. 

The  bluefish,  which  commonly  attains  a  weight  of  10  pounds  (the 
record  specimen,  taken  at  Nantucket  Island,  weighed  27  pounds),  has 
quite  a  reputation  for  the  size  of  its  appetite  and  the  ferocity  of  its 
attacks  on  schooling  menhaden,  mackerel,  herring,  or  other  small  fishes. 
According  to  old  and  often  quoted  accounts,  these  attacks  go  far  beyond 
what  is  necessary  to  satisfy  the  appetites  of  these  bold  sea  hunters,  which 
leave  a  trail  of  mutilated  fish  in  their  wake.  Writers  of  past  generations 
used  to  speculate  about  the  number  of  fish  destroyed  by  bluefish  each 
year,  seldom  agreeing  in  their  estimates  but  always  arriving  at  astronom- 
ical figures. 

Of  all  the  fish  tribe,  the  closest  relative  of  the  bluefish  that  is  widely 
known  is  the  pompano,  generally  voted  the  choicest  food  fish  of  the 
Atlantic  coast.  Like  its  relative,  the  bluefish  is  excellent  eating.  Since 
market  sizes  run  from  one  to  seven  pounds,  it  is  sold  whole  and  may  be 
baked,  broiled,  or  fried,  according  to  size  and  the  preference  of  the  cook. 

In  Florida,  where  the  only  important  commercial  catches  are  made, 
bluefish  are  taken  usually  between  mid-December  and  mid-February. 
The  same  gill  nets  that  take  the  Spanish  mackerel  also  capture  the  blue- 
fish, although  small  quantities  are  taken  also  by  trolling.  New  York  is 
an  important  market  for  these  southern  bluefish,  which  are  available  at 
a  season  when  the  fish  are  absent  from  northern  waters. 


16 


CONSERVATION    BULLETIN    3  7 


Red  Snapper  (Lutianus  hlackfordii) 

The  red  snapper  is  perhaps  the  most  widely  known  of  all  the  fishes  of 
the  Gulf  of  Mexico.  Although  the  annual  catch — 6  to  7  million  pounds 
— is  not  large  compared  with  the  major  fisheries  of  the  country,  the  red 
snapper  is  known  and  highly  regarded  in  many  cities  in  the  north,  to 
which  considerable  quantities  are  shipped.  It  is  usually  the  most  color- 
ful item  in  a  seafood  market's  display  case  or  window — a  large,  bril- 
liantly colored  fish  running  up  to  30  pounds  in  weight  and  2%  feet  in 
length.  Although  often  sold  whole,  one  of  the  larger  snappers  is  likely 
to  be  beyond  the  modern  family's  capacity  and  so  the  present  market 
forms  include  steaks  and  fillets,  quick  frozen  and  fresh.  Red  snapper  is 
available  throughout  the  year,  and  the  housewife  may  use  any  method 
of  preparation  she  likes,  for  it  is  good  broiled,  baked,  steamed,  or 
boiled.  The  meat  is  juicy,  white,  and  of  fine  flavor.  Most  people 
consider  a  baked  red  snapper  one  of  the  choicest  delicacies  the  sea 
provides. 


Figure   5.— The   Gulf   coast's   red   snapper   is   known   all    over   the    United   States   as   a    choice 

food    fish. 


Most  red  snappers  are  caught  in  the  Gulf,  with  the  most  extensive 
known  fishing  grounds — the  Campeche  3ank — lying  off  the  coast  of 
Yucatan  and  some  400  miles  from  Pensacola,  the  principal  United  States 
port  for  this  fish.  Small  boats  out  of  Galveston,  Mobile,  Panama  City, 
and  Tampa  also  fish  on  the  Campeche  Bank,  taking  both  snappers  and 
groupers,  which  commonly  occur  on  the  same  grounds.  There  is  also 
good  fishing  for  these  species  around  the  edges  of  smaller  banks,  known 
as  "lumps,"  off  the  United  States  coast  of  the  Gulf. 

At  least  five  species  of  snappers  are  taken  by  United  States  fishermen, 
although  the  red  snapper,  Lutianus  blackfordii,  is  caught  in  much  the 
largest  quantity.  About  Key  West,  the  yellowtail,  Ocyurus  chrysurus, 
is  the  most  important  of  all  the  snappers  and  is  regarded  as  one  of  the 
choicest  of  the  local  food  fishes.  It  averages  about  a  pound  and  is  com- 
monly taken  on  the  outer  reefs.  The  gray  snapper,  L.  griseus,  is  an  even 
more  abundant  fish  about  Key  West  but  because  it  is  also  exceedingly 
wary  not  as  many  are  caught.  This  snapper  has  an  interesting  habit  of 
traveling  in  schools  that  may  number  thousands  of  individuals.  If  not 
disturbed,  such  a  school  may  hover  in  one  place  for  hours,  easily  seen 
through  the  clear  water.     The  gray  snapper  is  a  good  food  fish  and 


FISH    AND    SHELLFISH  17 

market  sizes  are  from  one-half  to  five  pounds.  In  commercial  import- 
ance, the  muttonfish  (L.  analis)  probably  ranks  next  to  the  yellowtail  in 
the  Key  West  area.  It  averages  about  3  pounds  and  is  an  excellent  food 
fish.  The  schoolmaster,  L.  apodus,  is  also  rather  common  in  southern 
Florida  but  its  commercial  value  is  less  than  that  of  the  other  species. 

Besides  the  species  named  many  other  members  of  the  snapper  family 
inhabit  the  waters  of  the  West  Indies,  and  these  are  among  the  most 
beautiful  food  fishes  of  the  tropics.  In  his  "American  Fishes, "  G.  Brown 
Goode,  who  first  recognized  and  named  the  red  snapper,  says  they  are 
"the  tanagers  and  grosbeaks  of  the  coral  reefs." 

It  is  an  interesting  fact  that  the  red  snapper  was  not  named  until 
1878,  although  it  had  been  a  favorite  in  southern  markets  long  before 
that,  and  had  even  been  introduced  in  the  markets  of  New  York,  Wash- 
ington, and  throughout  the  Mississippi  Valley.  In  1878  Dr.  Goode,  then 
Assistant  Secretary  of  the  Smithsonian  Institution,  studied  a  number  of 
specimens  obtained  from  Florida  and  decided  the  fish  was  a  species  new 
to  science.  He  named  it  Lutjanus  black f or dii:  hwtjanus  (later  spelled 
Lutianus)  from  an  Asiatic  name  for  a  kindred  species;  blackfordii  in 
honor  of  Eugene  G.  Blackford,  then  Commissioner  of  Fisheries  of  New 
York  and  an  enthusiastic  ichthyologist. 

Strangely  enough,  very  little  is  known  about  the  life  history  of  the 
red  snapper,  in  spite  of  the  high  esteem  in  which  it  is  held.  According 
to  fishermen,  snappers  move  into  comparatively  shallow  water  in  the 
summer  and  in  the  fall  move  offshore  again.  At  all  seasons  they  seem  to 
shift  from  one  spot  to  another  according  to  food  supplies.  They  are 
believed  to  spawn  in  deep  water,  during  the  late  summer  and  fall. 

Although  the  young  of  related  species  commonly  appear  along  our 
southern  shores  and  may  be  caught  in  beach  seines,  young  red  snappers 
are  seldom  if  ever  taken  in  such  localities. 

Offshore,  the  red  snappers  live  in  deep  holes  and  rocky  gullies  where 
small  fish,  crabs,  shrimps  and  other  marine  creatures  which  are  their 
prey  are  abundant.  Usually  found  on  the  same  grounds  with  them  are 
the  groupers,  which  ordinarily  outnumber  the  snappers  about  two  or 
three  to  one.  They  prefer  warm  water  and  if  the  temperature  falls 
below  50°  they  are  likely  to  move  off  in  search  of  a  more  congenial 
climate.  Their  only  enemies  are  sharks  and  a  few  of  the  larger  carniv- 
orous fishes,  such  as  the  enormous  jewfish. 

The  center  of  the  red  snapper  fishery  is  now  Pensacola.  From  that 
port,  a  round  trip  to  the  Campeche  Bank  takes  about  23  days:  4  days 
each  for  the  trip  out  and  returning,  15  days  fishing.  Many  of  the  red 
snapper  boats  are  small  craft  but  owing  to  the  skill  and  experience  of 
their  skippers  and  crews  they  are  able  to  weather  the  heavy  blows  and 
rough  seas  of  the  Gulf. 

Snappers  are  taken  in  water  several  hundred  feet  deep,  and  most  of  the 
catch  is  made  with  hook  and  line.  According  to  Jarvis  (1935)  a  crew  of 
9  men  can  haul  in  from  7,000  to  10,000  pounds  a  day,  depending  on  the 
depth  of  the  water,  the  tide,  and  the  wind.  The  large  schooners  which 
fish  on  the  Campeche  Bank  take  most  of  their  catch  between  October 
and  April,  while  the  smaller  boats  fishing  along  the  United  States  coast, 
being  more  dependent  on  good  weather,  operate  largely  during  the 
spring  and  summer. 

Groupers  (Genera  Epinephelus  and  Mycteroperca) 

The  groupers  belong  to  one  of  the  largest  and  most  widely  distributed 
families  of  fishes  in  the  world — the  sea  basses  or  Serranidae.    The  sea 


18 


CONSERVATION    BULLETIN    3  7 


bass  tribe  is  so  numerous,  affects  other  fish  so  directly  because  all  its 
members  are  carnivorous,  and  is  found  in  so  many  locations,  that  it  is 
usually  considered  the  dominant  group  of  marine  fishes.  The  larger 
members  of  the  family  are  prized  as  food  wherever  they  occur. 

As  might  be  expected  in  such  a  large  family,  many  rather  diverse 
fishes  are  included  in  it.  The  sea  basses,  groupers,  jewfishes,  and  others 
belong  to  the  group,  which  numbers  about  120  species  in  North  and 
Central  America. 

Common  names  of  members  of  the  sea  bass  family  range  all  the  way 
from  gag,  scamp,  and  tally-wag  to  such  elegant  designations  as  the 
princess  rockfish.  The  name  grouper  is  thought  to  be  a  corruption  of 
the  Portuguese  "garoupa"  for  a  similar  species. 


Figure  6.— The   red   grouper  is  one  of  the  stople  food   fishes  of  the  south.     The   five  or  six 
million  pound  catch  does  not  permit  large  shipments  to  other  areas. 


Probably  the  most  abundant  of  the  groupers,  and  so  the  most  im- 
portant commercially,  is  the  red  grouper,  Epinephelus  morio,  found  from 
Virginia  to  Rio  de  Janeiro,  and  sometimes  straggling. northward  as  far 
as  Woods  Hole,  Mass.  It  grows  to  a  length  of  2  or  3  feet  and  may 
weigh  from  20  to  40  pounds.  Also  of  some  importance  commercially  are 
the  Nassau  grouper,  E.  striatus,  common  from  Florida  to  Brazil  and 
occasionally  taken  as  far  north  as  North  Carolina,  and  three  members  of 
the  genus  Mycteroperca — the  yellowfish  grouper,  M.  venenosa,  the 
black  grouper,  M.  bonaci,  and  the  gag,  M.  microlepis.  All  of  these  are 
found  in  greatest  abundance  on  the  Gulf  coast  but  most  also  occur  along 
the  South  Atlantic  coast  and  both  red  and  black  groupers  are  reported 
from  Woods  Hole.  Whether  these  are  fish  that  have  traveled  north  in 
the  Gulf  Stream  or  are  permanent  residents  of  New  England  is  some- 
thing that  has  not  been  determined. 

The  red  grouper  is  commonly  taken  on  the  same  grounds  with  the  red 
snapper,  and  is  caught  with  the  same  kind  of  gear.  On  some  grounds 
the  groupers  outnumber  the  snappers  by  two  or  three  to  one,  although 
they  are  rather  solitary  fish  and  seldom  school. 

Most  groupers  are  masters  of  the  art  of  camouflage.  When  resting 
among  corals  or  seaweeds,  where  there  is  a  background  of  mingled  lights 
and  shadows,  their  color  pattern  is  usually  banded.  This  banding,  as  a 
rule,  fades  out  as  soon  as  the  fish  rises  from  the  bottom  and  begins  to 
swim  through  open  water.  Then  it  usually  is  a  very  pale  and  ghostlike 
color  which  makes  its  outlines  hard  to  discern.    Apparently  the  changes 


FISH    AND    SHELLFISH  19 

are  not  related  to  motion  or  inactivity  alone ;  groupers  have  been  seen  to 
flash  their  color  bands  on  and  off  at  the  approach  of  another  fish  of 
different  species. 

For  these  interesting  observations  on  camouflage  as  well  as  on  many 
curious  habits  of  groupers  and  other  fishes  of  the  coral  reefs  we  are 
chiefly  indebted  to  the  late  Dr.  William  Longley.  During  many  hours 
spent  in  a  diving  helmet,  he  studied  fishes  in  their  natural  surroundings 
and  his  accounts  (Longley  and  Hildebrand,  1941)  of  what  he  saw  made 
the  sport  of  fish  watching  seem  at  least  as  interesting  as  bird  watching, 
and  nearly  as  practicable.  Some  groupers,  on  first  acquaintance,  would 
take  food  from  his  hand,  while  others  were  more  shy.  Red  groupers,  he 
found,  would  sometimes  allow  him  to  touch  them  even  though  he  offered 
them  no  food.  He  describes  one  of  the  bolder  fish — a  Nassau  grouper — 
as  follows: 

One  that  I  fed  frequently  usually  appeared  promptly  upon  my  approach  from 
under  the  coral  heads  and  got  under  foot  and  picked  at  my  pockets,  in  which  I 
carried  crawfish  tails  for  it.  It  could  scarcely  be  driven  far  enough  away  to  photo- 
graph, and  might  easily  have  been  captured  with  bare  hands  if  it  had  not  been 
so  strong,  hard,  and  slippery. 

One  of  the  grouper  family,  the  spotted  jewfish,  Promicrops  itaiara,  is 
one  of  the  largest  of  all  fishes,  the  record  specimen  weighing  693  pounds. 
Small  jewfish  taken  in  shallow  water  weigh  up  to  10  pounds  but  the 
average  weight  of  those  caught  commercially  about  Key  West  is  about 
125  pounds.  The  jewfish  is  caught  on  strong  tackle  resembling  that 
used  for  sharks  and  is  often  brought  in  alive  to  the  dock,  where  it  is 
hitched  to  a  stake  in  shallow  water  to  await  disposal  in  the  markets. 
When  the  fish  are  dressed  they  are  usually  cut  into  steaks,  and  the  local 
demand  is  so  heavy  that  only  limited  quantities  are  available  for  other 
markets.  The  spotted  jewfish  is  confined  to  tropical  waters  but  is  found 
on  both  Pacific  and  Atlantic  coasts. 

Most  of  the  groupers  when  fully  grown  weigh  as  much  as  40  or  50 
pounds,  although  market  forms  usually  range  from  5  to  15  pounds. 
They  are  sold  whole,  or  as  steaks  or  fillets.  They  are  usually  fried  or 
broiled,  or  the  fillets  may  be  baked  in  a  sauce.  Groupers  are  caught 
throughout  the  year.  The  chief  market  for  groupers  is  in  the  south- 
eastern States,  which  receive  about  three  fourths  of  the  catch.  Outside 
this  area,  St.  Louis,  Cincinnati,  and  Chicago  receive  a  limited  quantity. 

Sea  Trouts  (Genus  Cynoscion) 

The  sea  trouts  belong  to  the  croaker  family  and  so  are  related  to  the 
drums,  croakers,  and  spots  rather  than  to  the  well-known  fresh  water 
trout.  They  are  exclusively  marine  fishes  and  one  or  more  species  are 
found  all  along  our  coast  from  Cape  Cod  to  Texas.  The  spotted  sea  trout, 
Cynoscion  nebulosis,  is  the  most  important  of  the  group  in  the  southern 
fisheries,  being  taken  in  some  abundance  from  North  Carolina  south  with 
the  center  of  the  fishery  in  the  Gulf.  The  gray  sea  trout,  C.  regalis,  is 
taken  in  greater  quantity  than  the  spotted  in  the  fisheries  of  North 
Carolina,  but  farther  south  it  is  relatively  scarce  and  in  the  Gulf  it 
occurs  only  occasionally.  A  third  species  of  some  importance  is  the 
white  or  sand  sea  trout,  C.  arenarias,  found  only  in  the  Gulf.  Perhaps 
because  it  is  small — 11  to  15  inches — it  is  rather  neglected  by  fishermen. 

The  spotted  sea  trout  is  an  excellent  game  and  food  fish  that  brings  a 
good  price  at  all  seasons  of  the  year  and  is  universally  popular  in  the 
markets.  The  flesh  is  white,  tender,  and  of  delicate  flavor.  The  smaller 
sizes  are  sold  whole  and  are  best  broiled  or  fried.    The  larger  sizes  may 


20 


CONSERVATION    BULLETIN    3  7 


be  baked,  although  they  are  often  filleted  and  in  this  form  may  be 
broiled.  They  are  taken  throughout  the  year,  but  in  greatest  quantity 
in  spring,  summer,  and  fall. 

The  white  sea  trout,  the  smaller  species,  is  a  good  pan  fish.  Although 
it  is  taken  now  in  small  quantities — about  a  quarter  of  a  million  pounds 
compared  with  seven  million  pounds  of  the  spotted  sea  trout — probably 
it  will  receive  proper  appreciation  in  the  future  when  the  larger  species 
are  no  longer  able  to  meet  the  demand. 

At  least  in  some  areas,  the  spotted  trout,  unlike  most  shore  fishes, 
remains  in  the  shallow  waters  inshore  throughout  the  year,  instead  of 
moving  out  into  deeper  water  to  escape  the  cold.  In  North  Carolina  it 
is  one  of  the  few  fishes  available  to  the  haul  seine  fishermen  during  the 


Figure  7.— The  spotted  sea  trout  is  the  most  abundant  member  of  its  family  from  the  Caro- 
lines  to   Texas. 


lean  winter  season.  Ordinary  winter  temperatures  do  not  affect  the 
spotted  trout  adversely,  but  a  period  of  unusual  cold  will  cause  large 
numbers  of  them  to  become  numb  and  helpless.  At  such  times  they 
can  be  picked  up  by  the  bushel  along  the  shores.  If  taken  into  a  heated 
room  they  quickly  revive,  so  the  numbing  apparently  is  not  necessarily 
fatal. 

The  larger  trout  travel  in  small  schools,  preying  on  shrimps  and  small 
fishes.  They  themselves  are  taken  by  large  predatory  fishes  and  some- 
times are  followed  and  harried  by  schools  of  porpoises.  Unlike  the 
mullet,  famous  jumpers,  they  swim  low  in  the  water,  making  little  or  no 
ripple  at  the  surface.  When  close  to  the  shore  line,  they  usually  move 
in  with  the  rising  tide  and  offshore  with  the  ebb. 

In  contrast  to  the  red  and  black  drums,  the  spotted  trout  in  Texas 
usually  spawns  in  inshore  bays  and  lagoons,  rather  than  in  the  open 
Gulf.  The  spawning  season  extends  from  early  April  to  September  in 
Texas,  in  North  Carolina  probably  from  May  into  August.  The 
spawning  female  produces  from  half  a  million  to  a  million  eggs.  Prob- 
ably most  spotted  trout  mature  and  spawn  for  the  first  time  when  they 
are  two  years  old,  and  about  a  foot  long.  As  not  infrequently  happens 
among  fishes,  the  males  are  smaller  than  the  females,  the  average  dif- 
ference being  about  2  inches. 

In  Texas  most  of  the  commercial  catch  is  made  with  hook  and  line  in 
the  passes  or  channels  and  with  drag  seines  in  the  bays  and  lagoons.  A 
small  amount  of  gill-net  and  trammel-net  fishing  is  done  during  the- 
winter  along  the  northern  Texas  coast,  and  a  few  fish  are  taken  with 
light  surf  seines  along  the  beaches  of  the  Gulf  of  Mexico.    The  largest 


FISH    AND    SHELLFISH 


21 


catches  of  the  spotted  trout  are  made  on  the  west  coast  of  Florida  and 
in  Texas  and  North  Carolina. 

Redfish  or  Red  Drum  (Sciaenops  oceUata) 

The  redfish  is  really  silver  with  a  suffused  gleam  of  red  over  its  scales, 
like  a  color  reflected  on  metal.  Much  of  this  illusive  color  disappears 
soon  after  the  fish  is  taken  from  the  water,  so  that  the  redfish  in  the 
market  lacks  the  splendid  coloration  of  the  red  snapper  or  the  New 
England  rosefish.  It  is,  however,  a  graceful  and  handsome  fish,  longer 
and  more  slender  than  the  related  black  drum,  and  bearing  one  or  more 
conspicuous  black  spots  at  the  base  of  the  tail  fin. 


Figure  8.— The  redfish  is  a   member  of  the  drum  family,   but  in  this  species   only  the   males 
are  equipped  to  make  the  characteristic  drumming  noises,  while  the  females  are  silent. 


As  might  be  expected  of  a  fish  that  is  found  in  commercial  quantities 
all  the  way  from  New  Jersey  to  Texas,  the  redfish  is  known  by  various 
names.  Usually  called  redfish  on  the  Gulf  coast  where  it  is  most 
abundant,  it  is  known  as  the  red  drum  or  spot  bass  on  the  Atlantic  coast. 
Surf  anglers,  who  ask  for  nothing  better  than  a  struggle  with  this  gamy 
fish  in  the  tumbling  breakers,  often  call  it  the  channel  bass. 

For  market  purposes,  redfish  from  2  to  25  pounds  or  so  are  taken,  the 
larger  sizes  being  dressed  as  steaks  or  fillets.  Redfish  larger  than  25 
pounds  are  of  interest  chiefly  to  sportsmen,  being  less  tender  and 
palatable  than  younger  fish  and  also  being  protected  in  some  States  as  a 
spawning  reserve.  The  largest  specimens  on  record  were  about  five  feet 
long  and  weighed  about  75  pounds. 

The  redfish  is  one  of  the  more  important  food  fishes  of  the  Gulf  coast 
and  is  taken  in  some  quantity  also  along  the  South  Atlantic  shore.  The 
total  catch  is  about  a  million  and  a  half  pounds,  of  which  slightly  more 
than  half  is  caught  in  Florida.  Most  of  the  Florida  catch,  in  turn,  is 
taken  on  the  west  coast.  In  Texas,  where  the  redfish  ranks  fourth  in 
volume  of  catch  among  the  fin  fishes,  about  a  quarter  of  a  million 
pounds  are  caught.  The  spotted  sea  trout  is  the  only  fish  taken  in  larger 
quantity  in  Louisiana.  While  the  fishing  seasons  for  redfish  extend 
throughout  the  year  in  the  various  parts  of  the  southern  coast,  produc- 
tion peaks  come  in  the  period  from  February  to  June  and  again  in  the 
fall  in  Texas,  and  from  November  to  April  in  Florida. 

Although  the  redfish  has  been  known  to  fishermen  for  at  least  60  or 
70  years,  much  of  its  life  history  has  been,  and  still  remains,  a  mystery. 


22  CONSERVATION    BULLETIN    3  7 

Pearson,  (1928),  studying  the  species  on  the  southern  Texas  coast,  found 
that  the  fish  spawn  in  that  area  mainly  in  the  fall  of  the  year.  This 
part  of  the  coast  is  bordered  by  a  chain  of  long,  narrow  barrier  islands 
which  lie  off  the  mainland,  separated  from  it  by  a  series  of  bays.  Several 
channels  or  passes  connect  the  bays  with  the  open  Gulf,  and  Pearson 
found  that  the  redfish  tended  to  spawn  to  seaward  of  these  passes,  the 
newly  hatched  young  then  drifting  in  with  the  tides  and  currents  to  the 
bays  and  lagoons.  In  these  protected  waters  they  develop  rapidly  and 
soon  reach  a  stage  where  they  can  swim  under  their  own  power  and 
without  regard  to  prevailing  currents.  They  scatter  widely,  usually 
seeking  the  quiet,  grassy  bottoms  where  they  find  better  protection 
from  enemies  than  on  bare,  sandy'bottom.  Sometimes  this  habit  reacts 
to  their  destruction,  for  abnormally  low  tides  may  occasionally  drain 
the  water  out  of  these  shallow  flats  and  leave  the  young  fish  stranded  by 
the  thousand. 

With  the  coming  of  the  cold  weather  of  their  first  winter,  the  young 
redfish,  by  this  time  from  2  to  6  inches  long,  find  their  way  into  the 
deeper  bayous,  which  are  inland  channels  between  bays  or  running  into 
the  mainland  from  the  bays.  When  spring  comes  they  leave  the  inland 
waters  for  the  first  time,  some  going  out  through  the  passes  to  the  open 
waters  of  the  Gulf,  others  wandering  through  the  inland  bays  and 
lagoons.  Redfish  less  than  a  year  old  have  been  taken  75  miles  from 
the  nearest  pass,  through  which  they  must  have  been  carried  as  eggs  or 
larval  fish. 

After  their  first  year  most  redfish  winter  in  the  deeper  bays  or  in  the 
Gulf,  returning  to  inside  waters  in  the  spring.  The  fall  exodus  is  not 
especially  noticeable,  but  the  spring  migration  is  undertaken  by  large 
bodies  of  fish  moving  together  through  the  various  passes,  where  a  good 
many  are  taken  by  hook  and  line  fishing. 

The  adult  or  bull  redfish  wander  up  and  down  the  sandy  shores  of  the 
Gulf  throughout  much  of  the  year,  assembling  in  the  fall  about  the 
mouths  of  the  passes  to  spawn.  These  large  fish  are  seldom  seen  in  the 
bays  and  lagoons,  probably  because  they  are  easily  frightened  by  the 
disturbances  made  by  speedboats  in  the  shallow  waters,  Pearson  suggests. 

Redfish  are  themselves  no  mean  connoisseurs  of  seafood.  Shrimp  and 
crabs  are  their  favorite  foods.  They  also  eat  a  few  fish,  especially  mullet 
and  small  minnow-like  forms,  and  on  occasion  take  almost  any  of  the 
small  invertebrate  sea  life  which  they  come  upon  in  their  wanderings. 

Redfish  grow  very  rapidly.  They  are  about  13V2  inches  long  when  a 
year  old  and  about  21  inches  by  the  end  of  the  second  year.  It  is  be- 
lieved that  as  a  practical  conservation  measure,  they  should  be  pro- 
tected from  capture  until  they  are  about  16  inches  long,  or  approxi- 
mately a  year  and  a  half  old.  At  this  length  they  weigh  about  three 
times  as  much  as  at  12  inches,  and  are  a  more  desirable  market  size. 

Black  Drum  (Pogonias  cromis) 

The  black  drum  may  be  taken  anywhere  from  the  shores  of  Long 
Island  to  the  mouth  of  the  Rio  Grande,  but  it  is  only  in  Texas  that  it 
becomes  a  really  important  commercial  species.  In  that  State  it  ranks 
third  among  the  commercial  fisheries,  only  the  red  snapper  and  the 
spotted  sea  trout  being  taken  in  greater  quantity.  The  amount  of  drum 
caught  by  all  United  States  fishermen  during  recent  years  has  run  from 
one  to  two  million  pounds  annually,  and  of  this  Texas  furnishes  from  50 
to  80  percent.  Florida,  Louisiana,  and  North  Carolina  are  next  in  order, 
and  small  amounts  are  taken  in  all  the  other  southern  States. 


FISH    AND    SHELLFISH 


23 


The  drum  is  a  heavyweight  among  fishes — the  largest  specimen  known 
weighed  146  pounds — but  for  market  purposes  the  smaller  sizes  ranging 
from  8  to  20  inches  long  are  most  desired.  These  are  good  food  fishes 
and  lend  themselves  to  broiling  or  to  baking,  provided  fat  is  added,  for 
the  flesh  is  somewhat  lean. 

Quite  aside  from  its  food  qualities,  the  black  drum  is  an  exceptionally 
interesting  fish.  It  is  probably  the  best  musician  of  all  the  large  family 
of  drums  or  croakers  to  which  it  belongs.  People  who  have  always  sup- 
posed fish  to  be  universally  silent  creatures  are  surprised  to  learn  that 
many  of  them  have  voices  so  loud  that  they  can  actually  be  heard  a 
considerable  distance  above  the  surface  of  the  water.  These  vocal  ef- 
fects, in  the  case  of  the  drums,  are  produced  by  the  vibration  of  special 


Figure   9.— The   black    drum    is    sometimes    called    the    sea    drum    to    distinguish    it   from    its 
fresh-water   relative,  the   sheepshead   or   gaspergou. 


band-like  muscles  against  the  taut  air  bladder.  The  black  drum  is  now 
believed  to  be  the  subject  of  the  old  Indian  legends  in  the  country  about 
Pascagoula  Bay.  According  to  these  tales,  mysterious  music  of  sup- 
posedly supernatural  origin,  described  as  sweet,  plaintive,  and  low, 
could  be  heard  on  summer  evenings  apparently  issuing  from  the  water. 
Fishermen  are  now  well  acquainted  with  these  sounds,  and  know  that 
they  indicate  the  presence  of  a  large  school  of  drums.  Whether  the 
drumming  is  a  call  to  attract  the  opposite  sex  or  an  expression  of  con- 
tentment, as  at  feeding  time,  is  uncertain,  but  some  observers  think  the 
fish  are  most  vocal  during  their  spawning  migration.  The  males  are 
said  to  drum  very  loudly,  while  the  females  have  softer  voices. 

The  black  drum  has  such  large  and  competent  pharyngeal  teeth  that 
it  is  not  hard  to  guess  that  its  favorite  food  is  shellfish.  While  generally 
reported  to  be  an  oyster  pirate  of  no  small  capacity,  in  Texas  it  actually 
prefers  a  small  clam  which  is  very  abundant  in  the  shallow  bays  of 
that  State.  These  clam  beds  are  well  known  to  the  fishermen  as  yielding- 
good  hauls  of  drum.  The  fish  suck  the  clams  up  out  of  the  mud  in  which 
they  lie  and  crush  the  shells ;  although  they  swallow  so  many  shell  frag- 
ments along  with  the  meats  that  at  least  two  pounds  of  them  have  been 
taken  from  the  stomach  of  a  large  fish.  In  some  places  in  Louisiana 
stockades  or  fences  have  been  built  to  keep  the  drums  off  the  oyster  beds. 
They  apparently  are  great  gluttons,  and  when  feeding  stand  on  their 


24  CONSERVATION    BULLETIN    3  7 

heads  with  their  tails  out  of  the  shallow  water,  oblivious  to  all  but  the 
meal  at  hand. 

On  the  Texas  coast  the  drums  spend  a  great  deal  of  their  time  in  the 
shallow  inshore  bays,  both  summer  and  winter.  Sometimes  the  water 
in  the  passages  to  the  deeper  bays  becomes  so  shallow  that  they  have 
great  difficulty  navigating  these  channels  and  often  are  badly  lacerated 
by  passing  over  the  oyster  reefs.  It  is  said  that  farmers  used  to  chase 
the  large  drum  over  the  shallows  with  pitchforks. 

Young  drums  look  so  different  from  their  elders  that  they  were  for- 
merly supposed,  even  by  ichthyologists,  to  belong  to  a  different  species 
and  were  called  the  "banded  drum"  because  of  the  broad  brown  and 
white  bands  on  their  sides.  When  fully  grown  the  drum  is  a  silvery 
black  with  black  fins.  Those  taken  in  the  open  Gulf  are  often  more 
silvery;  those  in  inshore  waters  more  black.  The  whisker-like  barbels 
on  the  lower  jaw  are  typical  of  the  species. 

The  scales  of  the  black  drum  are  large  and  silvery  and  are  extensively 
used  in  making  ornamental  objects  and  jewelry.  They  are  so  hard  and 
so  firmly  embedded  that  it  is  necessary  to  remove  them  with  a  hatchet. 

Before  the  spawning  season — from  late  February  to  May — the  black 
drum  move  out  through  the  passes  into  the  Gulf,  spawning  later  near 
the  entrances.  The  young  drum  enter  the  inside  waters  soon  after 
hatching,  remaining  there  until  they  are  at  least  4  inches  long.  A  year- 
old  drum  is  about  10  inches  long;  a  five-year  fish,  23  inches.  The  fish 
spawn  when  about  two  years  old  (14  inches  long)  and  annually  there- 
after.   A  four-foot  drum  produces  about  6,000,000  eggs. 

Sportsmen  are  seldom  interested  in  the  black  drum,  which  is  a  more 
sluggish  fish  than  its  graceful  relative,  the  red  drum  or  redfish.  Some  of 
the  commercial  catch  is  made  on  hook  and  line,  but  the  heaviest  catches 
are  made  with  gill  nets  and  seines. 

Shrimp  (Penaeus  setiferus) 

The  Gulf  coast  is  the  chief  source  of  the  most  popular  crustacean 
in  the  United  States — the  shrimp.  The  American  people  eat  ten  times 
as  much  shrimp  as  lobster  and  half  again  as  much  as  they  consume 
of  crab.  And  of  the  150  million  pounds  of  shrimp  taken  from  the  sea 
each  year  for  cocktails,  salads,  and  appetizing  cooked  dishes,  85  per- 
cent comes  from  the  Gulf  coast  and  66  percent  from  the  State  of  Loui- 
siana alone. 

It  is  safe  to  assume  that  our  fathers  and  grandfathers,  unless  they 
happened  to  live  along  the  southern  coast,  knew  very  little  about 
shrimp,  for  in  the  1880's,  when  our  first  statistics  were  gathered,  the 
catch  was  only  about  10  million  pounds.  After  the  turn  of  the  century 
it  began  to  increase  slowly — 17  million  pounds  in  1905,  24  million  in 
1910,  57  million  in  1920.  By  1927  the  yield  of  this  small  but  delicious 
crustacean  had  passed  the  100-million-pound  mark,  and  in  1940  it 
reached  150  million  pounds. 

While  canned  shrimp  has  been  for  many  years  the  most  familiar 
market  form,  an  interesting  trend  toward  the  greater  use  of  fresh  and 
frozen  shrimp  set  in  during  the  period  of  the  war.  During  1943,  for 
example,  the  pack  of  canned  shrimp  on  the  Gulf  coast  declined  25  per- 
cent compared  with  the  previous  year,  although  the  catch  increased 
by  about  10  percent.  Although  the  immediate  reasons  for  the  decline 
of  shrimp  canning  are  the  result  of  war  conditions  such  as  the  shortage 
of  cannery  labor  and  the  high   prices   obtainable   for  fresh  shrimp, 


FISH    AND    SHELLFISH 


25 


many  dealers  predict  that  frozen  peeled  shrimp  will  be  the  mainstay 
of  the  postwar  industry. 

Since  some  shrimp  are  taken  throughout  the  year,  even  the  fresh 
product  is  available  at  any  season,  but  much  the  heaviest  production 
takes  place  in  the  late  summer  and  fall — usually  the  months  from 
August  through  December. 

In  life,  the  shrimp  looks  much  like  a  small  lobster.  However,  only 
the  meat  in  the  tail  or  abdomen  is  eaten,  for  the  claws,  which  in  the 
lobster  furnish  a  good  quantity  of  meat,  are  so  small  that  most  people 
would  fail  even  to  notice  them.  The  so-called  head  contains  the  vital 
organs  and  is  discarded  by  the  packer,  only  the  edible  portions  being 
shipped.     The  living  shrimp  is  a  pale  green  or  gray  creature  and  is 


Figure   10. — The  shrimp,   a   small   relative  of  the   lobster   and   the   crab,   supports  the   largest 
and   most  valuable  food  fishery  of  the  South. 

semi-transparent.     The  pink  or  reddish  color  develops  upon  cooking. 

The  life  story  of  an  individual  shrimp  begins  in  the  spring  or  sum- 
mer, when  spawning  takes  place  in  the  offshore  waters.  The  eggs, 
mere  specks  of  living  matter,  are  discharged  by  the  female  and  prob- 
ably remain  near  the  ocean  bottom  during  the  period  of  rapid  devel- 
opment that  precedes  hatching.  This  habit  of  the  shrimp  is  entirely 
different  from  that  of  the  closely  related  crabs  and  lobsters,  in  which 
the  females  carry  the  eggs  on  the  abdominal  appendages  until  they 
hatch. 

The  young  shrimp  hatch  from  the  eggs  in  a  very  short  time — prob- 
ably about  14  hours.  The  newly  hatched  larva  is  able  to  swim  about 
for  itself  and  probably  leaves  the  sea  bottom  very  soon  and  moves 
up  into  the  surface  waters.  As  it  floats  in  the  currents,  feeding  on 
the  minute  life  about  it,  the  baby  shrimp  undergoes  a  series  of  changes 
and  finally  comes  to  resemble  the  adult  form.  By  this  time  the  cur- 
rents have  carried  it  in  to  the  coast  line,  where  it  enters  the  shallow 
inside  waters  and  river  estuaries  and  again  takes  up  a  bottom-dwelling 
habit.  These  inside  waters  are  important  nursery  grounds  for  the 
young  shrimp  during  the  spring  and  summer. 

Since  the  spawning  season  of  the  shrimp  extends  over  a  period  of 
several  months — from  March  or  April  to  August  or  September — this 
means  that  some  young  shrimp  are  arriving  in  the  coastal  area  through- 
out the  entire  summer.  The  earliest  arrivals  reach  commercial  size 
during  the  month  of  July  and  by  September  practically  the  entire 
catch  is  made  up  of  young  shrimp  from  the  spawning  of  the  preced- 


26  CONSERVATION    BULLETIN    3  7 

ing  spring.  The  small  shrimp  winter  on  the  inside  grounds  and  along 
the  beaches,  but  most  of  the  larger  shrimp  move  out  into  the  open 
Atlantic  or  the  Gulf  to  escape  the  chilly  inside  waters.  Winter  fisheries 
for  these  large  shrimp  now  exist,  and  the  so-called  jumbo  shrimp  are 
taken  in  March,  April,  May,  and  June.  These  mature  shrimp, 
now  somewhat  more  than  a  year  old,  are  caught  throughout  the  summer 
on  the  offshore  fishing  grounds,  but  by  September,  when  a  new  crop 
appears  on  the  outside  grounds,  the  older  shrimp  make  up  a  very  small 
percentage  of  the  catch.  Presumably  the  life  span  of  the  shrimp  is 
between  one  and  two  years. 

Several  of  the  southern  States  have  adopted  measures  to  protect 
the  young,  rapidly  growing  shrimp  on  their  inshore  nursery  areas  dur- 
ing the  summer  and  fall.  By  waiting  until  the  shrimp  have  made  more 
of  their  growth,  fewer  individuals  would  be  required  to  make  up  the 
normal  150-million  pound  annual  catch. 

About  two-thirds  of  the  entire  United  States  catch  of  shrimp  is  taken 
in  Louisiana.  Texas  follows  with  ten  percent,  while  Mississippi,  Flor- 
ida, and  Georgia  each  contribute  5  or  6  percent,  North  Carolina  and 
Alabama  3  percent  each,  and  South  Carolina  only  1  percent.  Almost 
the  entire  catch  is  taken  by  small  otter-trawl  boats  which  drag  bag- 
like nets  along  the  bottom.  Cast  netting,  formerly  rather  important, 
still  brings  in  a  few  shrimp  and  very  small  incidental  catches  may  be 
made  by  other  gear  such  as  haul  seines. 

The  Oyster  (Ostrea  virginica) 

Southern  fishermen  make  more  money  from  oysters  than  from  any 
other  item  in  their  catch  except  shrimp  and  mullet.  The  annual  yield 
of  oysters  in  the  States  from  North  Carolina  to  Texas  is  some  20  mil- 
lion pounds,  from  which  fishermen  get  slightly  more  than  a  million 
dollars,  actually  a  higher  return  per  pound  than  they  receive  from  the 
more  productive  shrimp  and  mullet  fisheries. 

Although  the  production  of  oysters  now  is  about  the  same  as  fifty 
years  ago  considering  the  southern  area  as  a  whole,  every  State  from 
North  Carolina  south  shows  a  decline  with  the  single  exception  of 
Louisiana.  In  contrast,  that  State  produces  four  times  as  many  oysters 
now  as  it  did  in  the  1890's,  and  since  its  increase  about  matches  the 
decline  in  all  other  southern  coastal  States  combined,  the  total  output 
shows  little  change. 

The  oyster  that  grows  along  the  South  Atlantic  and  Gulf  coasts  is 
the  same  species  that  occurs  all  the  way  up  the  Atlantic  coast  to  Cape 
Cod,  and  sparingly  beyond  that  point  to  Prince  Edward  Island  in 
the  Gulf  of  St.  Lawrence.  It  is  one  of  three  species  of  oysters  taken 
commercially  in  the  United  States,  the  other  two  being  found  on  the 
Pacific  coast,  most  abundantly  in  the  State  of  Washington. 

In  point  of  size,  the  Atlantic  coast  or  eastern  oyster  is  midway  be- 
tween the  tiny  Olympia  oyster  of  Puget  Sound,  the  meat  of  which  is 
about  as  big  as  a  man's  thumb  nail,  and  the  enormous  Pacific  or 
Japanese  oyster,  which  has  been  known  to  grow  a  foot  or  more  in 
length,  although  it  averages  much  less.  Under  the  best  possible  condi- 
tions, the  eastern  oyster  might  attain  a  length  of  8  or  10  inches,  but 
actually  they  are  marketed  when  considerably  smaller.  Several  stand- 
ard market  sizes  of  shucked  oysters  have  been  established  by  custom, 
the  smallest  being  known  as  standards,  medium  size  as  selects,  and  the 
largest  as  counts. 

Most  oysters  are  now  sold  as  shucked  meats,  having  been  removed 


FISH    AND    SHELLFISH  27 

from  the  shells  in  shucking  houses  to  which  they  are  delivered  by  the 
tongers  or  dredgers.  Depending  on  their  size,  there  are  from  150  to 
300  eastern  oysters  in  a  gallon.  Although  the  trade  in  shelled  oysters 
is  on  the  wane,  some  oysters  are  still  shipped  in  the  shell,  the  demand 
for  this  product  coming  chiefly  from  hotels  and  restaurants.  Still 
others  are  canned.  Practically  all  of  the  canned  oysters  produced  in 
the  United  States  are  prepared  in  the  southern  area.  The  city  of 
Biloxi,  Miss.,  is  now  the  world  center  for  the  canning  of  oysters,  a 
title  once  held  by  Baltimore.  In  1943,  Mississippi  packed  85,151  cases 
and  Louisiana  79,614  cases,  the  remaining  50,090  cases  coming  from 
canneries  in  the  States  of  North  and  South  Carolina,  Georgia,  Florida, 
and  Alabama.  Normally,  some  oysters  are  canned  on  the  Pacific 
coast,  but  there  was  no  canning  in  this  area  in  1943  for  reasons  con- 
nected with  the  war. 

Oyster  cultivation,  which  reaches  its  greatest  development  in  New 
England  (Carson,  1943)  is  practiced  in  the  South  only  in  certain 
areas.  The  most  important  of  these  is  Louisiana,  where  practically 
all  oysters  come  from  cultivated  and  privately  leased  bottoms.  This 
practice  of  cultivation  has  kept  oyster  production  in  Louisiana  at  a 
high  level,  while  other  southern  States  have  declined.  There  are  in- 
dications that  scientific  oyster  farming  may  be  undertaken  in  North 
Carolina  and  some  additional  areas  after  the  war. 

Because  of  the  long  coastline,  oysters  grow  under  quite  a  variety 
of  conditions  between  Hatteras  and  the  southern  Texas  coast.  Many 
people  are  surprised  to  learn  that  oysters  never  grow  abundantly  on 
the  bottom  of  the  ocean  proper.  Instead,  they  thrive  best  in  enclosed 
bays,  sounds,  and  river  mouths,  where  the  salinity  has  been  reduced 
by  the  influx  of  fresh  water  from  rivers  and  streams.  In  the  more 
northern  bays  such  as  the  Chesapeake  or  the  Delaware,  they  may 
grow  some  distance  offshore  in  water  30  or  more  feet  deep,  but  along 
much  of  the  southern  coast  the  oyster  beds  are  mainly  between  low 
and  high  water  marks.  In  the  shallow  bays  and  estuaries  of  the  Gulf 
coast,  the  natural  oyster  beds  are  seldom  under  more  than  2  or  3  feet 
of  water.  In  southern  Florida,  oysters  sometimes  grow  attached  to 
the  aerial  roots  of  the  mangrove  trees,  submerged  only  by  the  high 
tide,  and  on  the  Gulf  coast  they  sometimes  grow  on  the  grass  at  the 
edges  of  the  marshes,  in  places  where  only  the  highest  tides  reach. 

On  the  Gulf  coast  the  spawning  season  of  the  oyster  is  a  very  long 
one,  beginning  in  March  and  continuing  into  November.  So  prolific 
is  the  oyster  that  a  female  may  produce  from  15  million  to  114  million 
eggs  at  one  spawning,  or  several  hundred  million  in  a  summer.  The 
eggs  and  sperms,  discharged  into  the  water,  unite  by  chance  and  the 
fertilized  egg  produces  a  free-swimming  larva  in  only  5  to  10  hours. 
The  young  oyster  swims  about  actively,  propelling  itself  by  the  vibra- 
tion of  minute,  hairlike  cilia.  For  about  two  weeks  it  is  a  creature 
of  the  tides  and  currents,  and  may  travel  far  from  the  place  where 
spawning  took  place.  At  the  end  of  this  period  it  is  a  fully  developed 
larva,  about  one  seventy-fifth  of  an  inch  in  diameter,  and  is  now  ready 
to  become  transformed  into  the  juvenile  form,  which  resembles  the  adult 
in  general  appearance.  Before  it  can  do  this,  it  must  give  up  its  free- 
dom of  motion  by  attaching  itself  firmly  (by  means  of  a  cementlike 
substance  which  it  secretes)  to  a  rock,  shell,  or  other  hard  object.  Once 
attached,  an  oyster  is  never  able  to  move  about  again  of  its  own  accord, 
although  oysters  growing  in  soft  mud  may  shift  their  position  by  brisk 
opening  and  closing  of  the  shells. 

The  act  of  setting  or  striking,  as  the  attachment  of  the  larval  oyster 


28  CONSERVATION    BULLETIN    3  7 

is  called,  is  full  of  danger,  for  even  a  few  grains  of  drifting  silt  are 
enough  to  smother  the  tiny  creature,  and  any  larvae  that  come  down 
on  a  surface  covered  with  algae  or  other  marine  growths  are  unable 
to  attach  themselves  and  quickly  die.  Those  which  make  a  successful 
landing  on  a  clean,  hard  surface  soon  change  from  the  larval  form  into 
miniatures  of  the  adult  oyster,  and  in  two  weeks  after  setting — at  the 
age  of  about  four  weeks — have  shells  a  quarter  of  an  inch  long.  Oysters 
from  this  size  to  two  inches  long  are  known  as  "seed  oysters."  In  south- 
ern waters,  where  the  water  ordinarily  is^warm  enough  for  oysters  to 
grow  and  feed  during  the  whole  year,  they  reach  market  size  in  two  to 
three  years.  About  four  years  are  required  in  Long  Island  Sound,  where 
the  cold  winters  cause  them  to  go  into  a  state  resembling  hibernation. 

On  the  uncultivated  reefs  of  the  South  are  found  many  long,  narrow, 
and  irregularly  shaped  oysters  which  are  known  as  coon  oysters,  a  name 
said  to  have  been  given  them  because  raccoons  feed  on  them.  These 
oysters  are  of  little  market  value  because  the  meats  are  small  and 
poorly  shaped.  The  coon  oyster  is  the  product  of  two  conditions  com- 
monly found  along  southern  coasts — soft,  muddy  bottoms  and  excessive 
crowding  of  oysters  in  a  narrow  zone  along  the  shore.  Single  oysters 
tend  to  sink  vertically  into  the  mud  so  that  the  shell  stands  on  one  end 
and  then  to  grow  into  long,  narrow,  sharp-edged  specimens.  Other 
oysters  crowd  in  closely  around  it,  attach  to  its  shells,  and  further  in- 
crease the  distortion  of  shape.  Small  coon  oysters  broken  apart  and 
planted  on  firm  bottom  change  their  form  and  develop  into  the  typical 
pear-like  shape  of  a  good  market  oyster.  Galtsoff  (1930)  points  out 
that  the  coon  oyster  is  not  a  geographic  race  but  a  product  of  its  en- 
vironment, and  says  that  in  earlier  days  typical  "coon  oysters"  existed 
in  Long  Island  Sound,  for  they  have  been  discovered  in  dredgings  at 
the  mouth  of  the  Housatonic  River. 

During  their  comparatively  short  lives  oysters  face  many  dangers. 
On  the  Gulf  coast,  especially,  flood  waters  from  the  rivers  or  the  break- 
ing of  levees  may  reduce  the  salinity  of  the  water  over  the  oyster  beds 
so  greatly  that  large  numbers  die.  Heavy  storms  may  bury  the  beds 
under  sand  or  dislodge  clusters  of  oysters  and  throw  them  on  the  beach. 
Gales  have  sometimes  been  known  to  establish  new  beds  by  shifting 
oysters  to  areas  where  none  had  grown  before.  Besides  storms  and 
floods,  oysters  also  have  many  natural  enemies.  Clams,  conchs,  and 
oyster  drills  which  bore  into  their  shells,  and  a  peculiar  sponge  of  a 
bright  yellow  color  which  bores  into  the  shell  and  causes  it  to  dissolve 
away  are  the  most  important  of  these  enemies  in  the  South. 

In  its  diet,  the  oyster  is  largely  vegetarian.  Most  of  its  food  con- 
sists of  the  minute,  one-celled  plants  called  diatoms  which  are  carried 
to  its  mouth  in  the  water  which  the  oyster  draws  through  its  gills.  The 
diatoms,  like  other  marine  plants,  are  nourished  by  the  various  minerals 
in  the  sea-water,  and  because  of  its  diet  the  oyster  itself  is  an  excellent 
source  of  copper,  iron,  iodine,  and  other  minerals  essential  in  human 
nutrition.  It  also  contains  most  of  the  essential  vitamins,  protein  of 
high  nutritive  value,  and  starch  in  the  easily  digested  form  known  as 
glycogen. 

Blue  Crab  (Callinectes  sapidus) 

The  common  blue  crab  of  the  South  Atlantic  and  Gulf  coasts  is  taken 
all  along  the  eastern  seaboard  from  New  York  to  Texas.  While  more 
blue  crabs  have  always  come  from  the  Chesapeake  Bay  than  from  any 
other  section,  the  yield  from  more  southern  areas  has  been  growing 
steadily.     The  State  of  Louisiana  now  produces  almost  half  as  many 


FISH    AND    SHELLFISH 


29 


crabs  as  are  taken  in  the  Chesapeake.  Compared  with  Louisiana's  14 
million  pounds,  North  and  South  Carolina  produce  about  4  million 
each,  Florida  about  6  million,  Georgia,  Alabama,  and  Mississippi  1  to 
2  million  each,  and  Texas  only  about  250,000  pounds. 

Unlike  its  relative,  the  shrimp,  the  blue  crab  is  only  occasionally 
found  in  the  open  sea  but  prefers  bays,  sounds,  and  the  mouths  of  rivers, 
where  it  lives  in  waters  ranging  from  true  ocean  saltiness  to  brackish 
or  even  fresh.  The  so-called  sweet-water  crabs  of  the  Atchafalaya 
River  are  taken  in  fresh  water  25  miles  from  its  mouth,  and  crabs  are 
common  in  the  Newport  River  of  North  Carolina.  However,  the  chief 
commercial  fisheries  are  in  the  bays  and  sounds,  where  the  crabs  are 
most  numerous. 


Figure  11.— The  blue  crab  of  the  Atlantic  and  Gulf  coasts  is  one  of  the  best  known  crustaceans 

in  the  world. 


Because  the  Chesapeake  fishery  for  crabs  has  been  longest  estab- 
lished and  also  because  the  bay  apparently  provides  ideal  surround- 
ings for  their  propagation,  the  habits  and  life  history  of  crabs  are  better 
known  in  the  Chesapeake  than  anywhere  else.  The  general  features  are 
probably  similar  in  all  sections,  however. 

Each  year  between  the  first  of  June  and  the  end  of  August,  in  the 
Chesapeake  Bay,  a  new  generation  of  crabs  is  produced.  At  this  sea- 
son the  female  extrudes  the  tiny  eggs,  each  about  1/100  inch  in  di- 
ameter, and  these  together  form  a  large  yellowish  mass  known  as  the 
"sponge"  and  remain  attached  to  the  female  crab.  As  the  young 
develop  within  the  eggs  the  sponge  becomes  darker.  They  hatch  in 
about  15  days.  Each  baby  crab  is  about  1/25  of  an  inch  long,  is  shaped 
somewhat  like  a  question  mark,  and  has  seven  pairs  of  appendages  and 
a  long  tail.  At  this  stage  it  is  konwn  as  a  "zoea."  As  it  grows  the 
zoea  sheds  its  shell  several  times.  After  the  fifth  or  sixth  shedding 
it  changes  into  a  form  that  is  more  truly  crablike.  This  is  the  "mega- 
lops."  Probably  the  first  molt  of  the  megalops  changes  the  developing 
youngster  into  the  true  crab  form,  but  it  is  decidedly  a  miniature  crab, 


30  CONSERVATION    BULLETIN    3  7 

only  1/8  inch  across.  This  stage  is  reached  about  one  month  after 
hatching. 

After  the  megalops  stage  the  crab  molts  about  15  times  before  reach- 
ing maturity,  at  first  every  6  days  and  then  after  gradually  lengthen- 
ing periods  up  to  about  25  days  between  final  molts.  Ordinarily  the 
crab  gains  about  one-third  in  size  with  every  molt.  Crabs  reach  their 
full  growth  and  maturity,  and  cease  to  molt,  during  their  second  sum- 
mer, when  12  to  14  months  old. 

For  the  female  crab,  mating  occurs  only  once  in  her  life — immediately 
after  the  last  molt,  when  she  is  still  in  the  soft-shelled  condition.  In 
the  Chesapeake  Bay  this  is  usually  during  the  last  of  July  or  August. 
The  sperm  received  from  the  male  at  this  time  is  retained  by  the  fe- 
male and  used  to  fertilize  such  successive  batches  of  eggs  as  she  may 
lay  during  the  remainder  of  her  life.  Usually  the  first  spawning  takes 
place  when  the  female  is  about  2  years  old,  or  during  the  summer  fol- 
lowing mating.  It  is  believed  that  some  females  live  over  another 
winter  and  deposit  more  eggs  when  3  years  old,  but  that  few  or  none 
live  longer  than  this.  The  life  span  of  the  male  is  probably  about  the 
same  length. 

In  the  Chesapeake  most  of  the  young  crabs  hatched  in  the  lower 
part  of  the  bay  soon  begin  a  northward  migration.  Cold  weather  inter- 
rupts this  journey  and  they  settle  to  the  bottom  and  cease  to  feed  and 
also  to  grow  until  conditions  are  more  to  their  liking.  In  the  spring 
their  migration  is  resumed,  growth  proceeds,  and  finally  they  reach 
Maryland  waters  as  nearly  mature  crabs.  It  follows  that  the  mating 
of  the  majority  of  the  blue  crabs  of  the  bay  takes  place  in  Maryland. 
After  mating  the  females  return  to  the  lower  bay,  but  most  of  the 
males  remain  behind,  spending  the  winter  in  deep  holes  or  in  creeks 
and  rivers.  Only  about  a  fifth  of  the  crabs  taken  in  the  lower  part  of 
the  bay  during  the  winter  are  males.  Nearly  all  the  sponge-bearing 
crabs  are  found  in  Virginia  waters. 

Many  people  have  the  mistaken  impression  that  the  so-called  "soft 
crab"  is  a  distinct  species.  The  term  refers  to  any  crab  which  has 
shed  its  old  shell,  before  the  new  shell  has  hardened.  As  the  soft- 
shelled  crab  is  considered  especially  good  eating,  large  numbers  of  young- 
crabs  are  sought  in  the  spring  and  early  summer  while  they  are  still 
molting. 

A  newly  molted  crab  is  particularly  helpless  to  defend  itself  or  even 
to  escape  by  swimming.  The  actual  casting  of  the  shell  requires  only 
a  few  minutes  if  all  goes  well — up  to  an  hour  if  the  crab  is  disturbed 
or  has  suffered  recent  injury.  In  a  series  of  convulsive  movements, 
the  crab  withdraws  its  five  pairs  of  legs  from  the  old  encasements, 
having  greatest  difficulty  with  the  large  claws,  and  backs  out  of  the  shell. 
Under  the  old  shell  the  skin  is  soft  and  wrinkled  and  the  animal  is 
flabby  and  helpless.  Ordinarily  the  crab  selects  some  sheltered  place 
to  undergo  the  molt,  for  during  the  12  hours  until  the  new  shell  hardens 
it  is  in  imminent  danger  of  being  set  upon  by  large  fish  or  even  by  its 
own  kind,  or  of  being  scooped  up  in  a  crabber's  dip  net. 

Crabs  have  the  curious  ability  to  cast  off  one  or  more  of  their  legs 
and  will  use  this  habit  to  escape  if  caught  and  held  by  one  leg.  A  sud- 
den drop  in  temperature  may  sometimes  cause  one  or  several  legs  to 
be  snapped  off.  The  break  always  occurs  across  one  of  the  joints,  which 
protects  the  crab  from  the  danger  of  bleeding  to  death.  Ordinarily,  the 
legs  are  regenerated.  As  many  as  three  or  four  replacements  of  the 
same  appendage  are  possible.  In  the  regenerative  process,  three  or 
four  molts  are  sufficient  to  provide  a  new,  full-size  limb. 


FISH    AND    SHELLFISH  31 

On  the  Gulf  coast  the  heaviest  production  falls  in  the  months  of 
May,  June,  July,  and  August.  Some  crabs  are  sold  whole  as  taken 
from  the  water,  in  which  case  they  should  be  alive  when  purchased, 
whether  in  the  hard  or  soft  shelled  condition.  Many  people,  however, 
prefer  to  buy  the  meat  ready  for  use,  and  so  most  of  the  crabs  caught 
on  the  Atlantic  and  Gulf  coasts  go  to  the  packing  houses  where  they 
are  cooked  and  the  meat  removed  and  packed  in  convenient  containers. 
Except  for  one  cannery  in  South  Carolina  and  two  in  Louisiana,  the 
canning  of  crabs  is  confined  to  the  Pacific  coast,  where  a  different 
species  is  taken,  and  to  a  small  canning  industry  for  the  rock  crab 
in  New  England. 

At  the  present  time  the  most  important  markets  for  fresh  crab  are 
the  cities  of  the  Atlantic  seaboard,  which  have  long  been  familiar  with 
this  perishable  product  and  know  how  to  give  it  the  special  handling  it 
requires.  On  the  other  hand,  few  mid-western  cities  know  the  blue 
crab  as  well,  and  as  a  result  the  interior  markets  to  which  the  Gulf 
coast  product  could  most  profitably  be  shipped  have  not  been  fully 
developed.  This  lack  of  near-by  markets  for  fresh  crab  has  been  an 
important  factor  in  holding  back  the  devolpment  of  a  large  fishery  in 
the  Gulf.  Whether  it  could  ever  become  as  large  as  the  Chesapeake 
fishery,  however,  is  something  that  cannot  be  predicted  until  the  crab 
populations  of  the  region  are  investigated  by  a  proper  biological  study. 

Other  Fish  and  Shellfish 

The  menhaden  is  one  of  the  most  interesting  fishes  on  the  Atlan- 
tic seaboard.  It  supports  one  of  the  oldest  fisheries  in  America,  ranks 
second  in  volume  of  production  a<mong  all  United  States  fishes,  and 
is  caught  in  greater  quantity  on  the  South  Atlantic  and  Gulf  coasts 
than  all  other  fin  fishes  combined.  Despite  its  importance,  it  is  one 
of  the  least  known  fishes  in  our  waters,  for  few  people  have  ever 
heard  of  it.  The  explanation  of  this  surprising  situation  is  simple. 
Although  almost  every  person  in  the  United  States  has  at  some  time 
been  indebted  to  the  menhaden  for  something  he  eats,  wears,  or  uses, 
the  fish  has  seldom  been  used  extensively  as  food  and  so  has  not  become 
familiar  to  the  public.  It  is  included  in  this  bulletin  on  food  fishes 
for  two  reasons:  the  recent  development  of  canned  menhaden  which 
may  make  this  product  generally  available  after  the  war,  and  the  fact 
that  meal  and  oil  derived  from  menhaden  are  so  important  to  our  live- 
stock and  poultry  industries  that  this  fish  is  only  one  step  removed 
from  the  ham,  fried  chicken,  and  eggs  on  all  our  tables. 

Although  the  fact  is  seldom  recognized,  several  species  of  menhaden 
are  found  in  United  States  waters,  the  most  important  one  on  the  Atlan- 
tic coast  being  Brevoortia  tyrannus.  The  menhadens  occur  all  the  way 
from  Nova  Scotia  to  Florida,  and  southward  to  Brazil,  but  are  seldom 
taken  north  of  Cape  Cod,  and  only  small  and  scattered  fisheries  are 
carried  on  in  the  Gulf  of  Mexico.  In  the  South  Atlantic  region  the  most 
important  fisheries  are  located  in  North  Carolina  and  Florida,  the  chief 
menhaden  center  being  the  town  of  Beaufort,  N.  C.  The  catch  for 
the  entire  area  amounts  to  about  250,000,000  pounds,  or  68  percent  of 
the  catch  of  all  species  of  fish,  and  43  percent  of  the  catch  of  all  fish 
and  shellfish  combined.  The  total  United  States  catch  of  menhaden 
is  about  600,000,000  pounds. 

Although  enterprising  Yankees  canned  menhaden  as  sardines  a  cen- 
tury ago,  there  had  been  little  use  of  this  fish  as  food — except  by  fisher- 
men or  other  people  in  fishing  communities — until  a  small  canning  indus- 


32 


CONSERVATION    BULLETIN    3  7 


try  was  developed  at  Fernandina,  Fla.,  during  the  war.  Practically 
the  entire  output  of  this  enterprise  has  been  sent  to  Great  Britain 
through  Lend-Lease  channels,  but  some  canning  has  also  been  done  at 
Reedville,  Va.,  and  this  product  has  found  its  way  into  the  grocery 
stores  of  nearby  States.  In  all  probability,  canned  menhaden  will  be- 
come familiar  to  a  much  larger  public  after  the  war  when  large-scale 
canning  becomes  feasible. 

Being  members  of  the  herring  family,  the  menhadens  resemble  the 
more  familiar  shad  and  the  river  herrings  or  alewives  in  appearance, 
and  general  characteristics.  Like  these  relatives,  they  have  a  rich 
and  oily  flesh.  Most  of  the  oil  is  removed  during  the  canning  process 
and  only  the  large,  meaty  cuts  from  the  back  region  are  packed. 


Figure   12.— The  menhaden,  a  member  of  the  herring  family,  swims  in  large  schools  at  the 

surface  of  the  sea. 


The  food  of  menhaden  consists  entirely  of  the  minute  surface  life 
of  the  sea  which  they  strain  from  the  water  by  means  of  a  sieve-like 
structure  in  the  back  of  the  mouth.  Large  schools  of  feeding  menhaden 
give  the  water  a  brassy  tinge  because  of  the  strong  yellow  luster  of 
their  flanks,  and  this  color  helps  fishermen  to  locate  the  schools. 

For  several  generations  the  manufacture  of  meal  and  oil  has  over- 
shadowed all  other  uses  for  menhaden.  One  third  of  all  the  fish  meals 
and  a  quarter  of  the  marine  animal  oils  produced  in  the  United  States 
are  products  of  the  menhaden  industry.  The  meals  are  fed  to  hogs 
and  poultry  to  provide  the  indispensable  animal  protein  in  their  diet; 
the  oils  are  used  in  preparing  fortified  vitamin  feeding  oils  for  poultry. 
The  industrial  uses  of  menhaden  oil  are  many.  It  is  a  constituent  of 
many  paints,  varnishes,  insect  sprays,  printing  inks,  and  soap.  The 
oil  is  used  also  as  a  lubricant  of  machinery,  in  aluminum  casting,  and 
in  leather  tanning. 

The  shad,  Alosa  sapidissima,  was  formerly  one  of  the  most  impor- 
tant food  fishes  of  the  South  Atlantic  coast,  but  the  catch  has  now 
declined  to  only  about  a  tenth  of  the  10,000,000-pound  level  at  the 
turn  of  the  century.  Despite  its  reduced  importance,  however,  it  still 
brings  the  fishermen  of  Georgia  a  larger  income  than  any  other  fin 
fish,  and  in  South  Carolina  it  is  second  in  value  only  to  the  mullet. 
North  Carolina,  with  an  annual  catch  of  about  800,000  pounds,  ranks 
fourth  among  the  States  of  the  Atlantic  seaboard  in  the  production 
of  shad. 

Like  the  salmon,  the  shad  is  a  fish  that  spends  most  of  its  time  in 
the  ocean  but  runs  into  coastal  rivers  to  spawn  above  tidewater.    The 


FISH    AND    SHELLFISH  33 

young  are  hatched  in  fresh  water  and  at  the  end  of  their  first  sum- 
mer drop  down  stream  and  enter  the  ocean.  They  are  not  seen  after 
they  leave  the  streams  until  they  return,  three  or  four  years  later,  as 
mature  adults. 

The  shad  is  a  seasonal  fish  all  along  the  coast,  and  is  caught  earlier 
in  Florida  than  anywhere  else.  The  runs  of  shad  begin  there  in  Decem- 
ber, and  the  first  shad  to  reach  northern  markets  come  from  Florida. 
The  largest  catches  in  that  State  are  made  in  the  St.  Johns  River,  and 
at  a  considerable  distance  above  its  mouth.  The  shad  runs  in  North 
Carolina  begin  in  February. 

The  shad  is  the  largest  and  most  prized  member  of  the  herring  family, 
and  is  generally  regarded  as  one  of  the  best  food  fishes  of  the  Atlantic 
coast.  In  the  1870's  it  was  transplanted  to  the  Pacific  coast,  where  it 
does  not  occur  naturally,  and  it  took  hold  so  well  in  its  new  surround- 
ings that  it  now  occurs  from  California  to  southeastern  Alaska,  and 
shipments  are  regularly  made  from  the  Pacific  coast  to  eastern  markets. 
The  decline  in  its  abundance  on  the  Atlantic  coast  is  the  result  of  ex- 


8,105 

Figure  13.— The  shad  may  again  be  abundant  in  many  southern  rivers  if  proper  conservation 

measures  are  applied. 

cessive  fishing  and,  in  some  waters,  to  pollution  or  to  dams  without 
adequate  fishways.  The  fact  that  a  complete  recovery  of  the  much 
depleted  Hudson  River  runs  has  been  brought  about  by  careful  regula- 
tion of  the  catch  indicates  that  in  the  future,  if  proper  management  prac- 
tices are  followed,  we  may  have  more  shad  all  along  the  Atlantic 
seaboard. 

Shad  is  ordinarily  sold  fresh,  although  some  is  frozen  in  the  Middle 
Atlantic  area.  Market  sizes  are  from  1%  to  8  pounds.  It  is  usually 
sold  whole,  and  a  baked,  stuffed  shad  is  considered  one  of  the  finest 
seafood  dishes.  A  very  choice  market  form,  to  be  recommended  for 
the  ease  of  preparation,  is  boned  shad  fillets. 

The  common  pompano,  Trachinotus  carolinus,  is  one  of  the 
choicest  of  all  food  fishes  although  the  supply,  unfortunately,  is  not 
large.  The  annual  catch  is  only  a  little  more  than  half  a  million 
pounds,  almost  all  of  which  is  taken  in  Florida.  Because  the  pompano 
is  so  highly  regarded  for  the  rich,  yet  peculiarly  delicate,  flavor  of  its 
flesh,  fishermen  receive  several  times  as  much  for  it,  per  pound,  as  they 
do  for  most  other  fishes.  The  height  of  the  pompano  season  is  from 
late  January  to  April.     The  average  market  size  is  about  1  to  l1/^ 


34 


CONSERVATION    BULLETIN     3  7 


pounds.  Most  of  the  catch  is  marketed  fresh,  and  large  shipments  are 
made  to  northern  markets. 

Adult  pompanos  are  found  from  Virginia  to  Brazil  and  the  young 
stray  as  far  north  as  Cape  Cod.  Except  for  the  wanderings  of  the 
young,  this  fish  does  not  seem  to  migrate  extensively. 

The  pompano  is  a  member  of  the  family  Carangidae,  which  includes 
the  amberfish,  the  jacks,  and  the  lookdown  fish.  It  is  a  thin  fish  with 
a  deeply  forked  tail  and  is  covered  with  small  scales  with  a  blue,  silvery, 
or  golden  luster.  It  is  toothless  in  the  adult  stage,  but  despite  this 
fact  feeds  on  small  mollusks  and  crustaceans.  Since  it  seldom  takes 
the  hook,  it  is  not  a  game  species. 

Not  a  great  deal  is  known  of  its  life  history.  It  seems  to  prefer 
sandy  bottom  near  shore  where  it  feeds.  Schools  of  pompano  are 
abundant  about  inlets,  where  they  play  in  and  out  with  the  tides.  In 
North  Carolina  small  ones,  from  half  an  inch  to  two  inches  long,  are 
often  seen  in  the  surf  on  the  outer  beaches,  the  waves  repeatedly  cast- 
ing them  out  on  the  wet  sand,  from  which  they  jump  back  into  the 
next  breaker. 


Figure  14.— Of  the  three  species  that  occur  on  the  Atlantic  coast,  only  the  common  pompano, 
shown  here,  is  an  important  food  fish. 


The  snook,  robalo,  or  sergeant  fish,  Centropomus  undecimalis, 
is  a  common  fish  of  the  southwest  coast  of  Florida,  where  it  supports 
a  small  commercial  fishery  and  is  also  one  of  the  most  important  sport 
fishes  of  the  region.  It  also  occurs  on  the  coast  of  Texas  and  south- 
ward through  the  West  Indies.  In  the  United  States  it  is  the  only  well 
known  representative  of  its  family,  a  group  of  tropical  fishes  found 
on  both  coasts  of  America. 

In  Florida  the  snook  is  taken  by  several  different  kinds  of  gear,  of 
which  hand  lines  are  the  most  important.  Large  fish  often  swim  close 
inshore,  where  most  of  the  fishing  is  done.  The  snook  also  ascends 
streams,  but  seldom  goes  above  the  limits  of  brackish  water. 

The  snook  is  .a  good  food  fish,  with  an  excellent  flavor  and  a  white 
and  flaky  meat.  The  annual  catch  runs  to  about  half  a  million  pounds. 
The  average  size  is  3  pounds,  though  a  large  snook  may  weigh  as  much 


FISH    AND    SHELLFISH  35 

as  30.     The  larger  fish  are  marketed  as  steaks,  and  are  best  baked  or 
broiled. 


Figure   15.— Called  Gulf  pike,  sergeant  fish,  brochet  de  mer,  and  snook,  this  is  by  any  name 
an  excellent  market  fish  and  a  favorite  of  sportsmen. 

The  grunts  make  up  a  large  family  of  fishes  which  are  related  to 
the  snappers  and  are  found  chiefly  in  tropical  waters.  Many  of  them 
are  important  food  fishes  wherever  they  are  locally  abundant.  One  of 
the  best  known  along  the  South  Atlantic  coast  is  the  pigfish,  Ortho- 
pristes  chrysopterus.  Fishermen  often  take  it  in  haul  seines,  for  it  is 
common  on  sandy  shores.  Averaging  about  half  a  pound — although 
some  weigh  as  much  as  two  pounds — the  pigfish  falls  in  the  panfish  cate- 
gory and  is  considered  very  good  eating. 

Several  grunts  of  the  genus  Haemulon  are  among  the  staple  food 
fishes  of  the  Key  West  area,  where,  according  to  the  colloquial  expres- 
sion, when  all  other  food  supplies  fail  the  natives  fall  back  on  "grits 
and  grunts."  The  white  and  yellow  grunts,  Haemulon  plumieri  and 
H.  sciurus,  are  the  most  abundant  there. 

Grunts  are  so  named  because  they  are  able  to  produce  as  audible 
sound,  although  scientists  disagree  as  to  just  how  this  is  done.  Some 
believe  it  is  by, scraping  the  pharyngeal  teeth  against  the  gill  arches, 
with  the  air  bladder  acting  as  a  resonator. 

Grunts  are  often  seen  in  shallow  water  and  about  docks,  where  they 
nose  into  cracks  and  crevices  in  search  of  food.  Schools  of  them  lie 
rather  quietly  during  the  day,  but  break  up  and  scatter  to  feed  at  night. 

Some  species  of  grunts  are  remarkable  in  having  the  inside  of  the 
mouth  colored  a  brilliant  red.  Just  what  purpose  this  color  serves  is 
uncertain,  but  two  fish  in  pugnacious  mood  will  sometimes  rush  at  each 
other  with  their  mouths  wide  open  and  the  brilliant  interiors  fully 
displayed. 

The  sheepshead,  Archosargus  probatocephalus,  of  the  Atlantic  and 
Gulf  coasts  might  be  seen  anywhere  from  Texas  to  the  Bay  of  Fundy, 
but  it  is  abundant  only  in  the  South  and  the  center  of  the  commercial 
fishery  is  on  the  west  coast  of  Florida.  The  annual  commercial  catch 
now  amounts  to  half  a  million  pounds  or  more,  although  formerly  it  was 
much  larger.  As  a  member  of  the  porgy  family  the  sheepshead  is  closely 
related  to  the  familiar  scup  or  porgy  that  is  an  important  commercial 
fish  northward  from  Virginia.  It  is  not  to  be  confused  with  the  fresh- 
water sheepshead,  which  is  a  member  of  the  drum  or  croaker  family. 

The  sheepshead  is  often  taken  about  wharves,  breakwaters,  and 
sunken  wrecks  where  it  finds  abundant  food.    It  is  fond  of  crabs,  oysters, 


36 


CONSERVATION    BULLETIN    3  7 


and  other  shellfish  which  it  crushes  easily  in  its  strong  teeth.  In  inlets, 
it  moves  in  and  out  with  the  tides  and  in  the  South  does  not  seem  to 
migrate  extensively.  During  the  spawning  season,  which  occurs  in  the 
spring,  the  sheepshead  assemble  in  schools  and  move  into  shallow  water 
to  deposit  the  eggs. 

The  largest  sheepshead  on  record  weighed  about  20  pounds  but  the 


Figure  16.— The  pigfish  is  one  of  the  commonest  food  fishes  along  South  Atlantic  shores. 

average  size  is,  of  course,  much  less.  The  larger  fish  may  be  filleted, 
the  smaller  sold  whole.  The  meat  is  white,  tender,  and  pleasantly 
flavored. 

A  related  species,  the  pinfish  Lagodon  rhomboides,  is  a  food  fish  of 
some  importance  in  Florida  and  North  Carolina. 


Figure  17.— Salt-water  anglers  as  well  as  commercial  fishermen  take  the  sheepshead,  a  fish 
that  prefers  warm  southern  waters. 


FISH    AND    SHELLFISH  37' 

The  blue  runner  or  hardtail,  Caranx  crysos,  has  always  been 
esteemed  as  a  food  fish  on  the  Gulf  coast,  where  it  seems  to  be  most 
abundant,  but  it  has  never  been  taken  in  large  quantities,  the  catch 
usually  falling  well  below  a  million  pounds.  Recently,  there  has  been 
some  increase  in  the  production  of  this  fish  and  in  1942  some  700,000 
pounds  were  taken  on  the  east  coast  of  Florida,  in  addition  to  the  Gulf 
coast  catch.  Too  little  is  known  about  the  biology  of  this  fish  to  predict 
how  large  a  fishery  it  could  support,  but  it  seems  likely  it  could  provide 
a  considerably  larger  poundage  than  is  now  being  taken. 

Like  the  pompano,  the  blue  runner  is  a  member  of  the  family  Carangi- 
dae,  all  of  which  are  active  swimmers  inhabiting  warm  seas.  Although 
found  all  the  way  from  Cape  Cod  to  Brazil,  the  blue  runner  is  taken 


Figure   18. — The  blue  runner  is  one  of  the  under-utilized  species  of  the  South. 

commercially  only  in  Florida  and  Alabama.  It  has  many  different 
common  names:  horse  crevalle  in  North  Carolina,  jurel,  skipjack,  and 
jack-fish  in  Florida. 

In  the  vicinity  of  Pensacola,  the  migratory  schools  of  the  blue  runner 
appear  in  April,  are  most  numerous  in  May.  Along  the  southern  coast 
of  Florida  they  are  abundant  in  winter.  They  follow  schools  of  small 
fish,  sometimes  leaping  out  of  the  water  in  pursuit  of  them.  They,  in 
their  turn,  are  chased  by  sharks  and  porpoises,  and  are  sometimes  driven 
ashore  in  numbers. 

The  blue  runner  is  a  slim  bodied  fish,  rather  more  green  than  blue 
above,  golden  yellow  or  silvery  below.  It  attains  a  length  of  20  inches 
and  a  weight  of  4  pounds.  Most  of  the  catch  is  taken  in  haul  seines, 
which  are  operated  in  shallow  water  along  the  shore. 

The  crevalle  or  common  jack,  Caranx  hippos,  a  close  relative  of 
the  blue  runner  and  the  pompano,  is  taken  in  quantities  of  about  a 
quarter  of  a  million  pounds  on  the  west  coast  of  Florida,  but  nowhere 
else  in  important  numbers.  This  is  the  most  abundant  of  the  various 
species  of  jacks  or  runners  that  are  found  about  Key  West.  It  is  a  good 
food  fish,  commanding  a  ready  sale  in  the  markets.  Most  of  the  com- 
mercial catch  is  taken  in  gill  nets,  but  the  crevalle  may  also  be  taken 
by  trolling  and  is  of  some  interest  to  anglers.  The  average  weight  is 
about  a  pound,  although  it  not  infrequently  runs  to  10  pounds  and  the 
record  weight  is  about  20.  Although  it  is  found  throughout  the  year,  it 
is  most  common  during  the  winter  months. 


38 


CONSERVATION    BULLETIN    3  7 


Figure  19.— The  crevalle,  which  occurs  on  the  eastern  seaboard  as  far  north  as  Massachusetts, 
has  a  number  of  common  names,  such  as  albacore,  horse  mackerel,  cavally,  and  jack. 

The  flounders  are  excellent  food  fishes,  with  white,  flaky,  and  del- 
icately flavored  meat,  but  the  total  catch  in  the  south  amounts  to  only 
about  a  million  pounds  by  commercial  fishermen.  While  some  are  taken 
in  each  of  the  coastal  States,  the  largest  catches  are  made  in  North 
Carolina,  Florida,  and  Texas. 

Three  kinds  of  flounders  are  taken  along  the  South  Atlantic  and  Gulf 
coasts,  although  one  of  them  is  so  small  that  it  has  little  commercial 
importance.  These  are  the  summer  flounder,  Paralichthys  dentatus, 
found  on  the  Atlantic  coast  as  far  south  as  the  northern  part  of  Florida ; 
the  southern  flounder,  P.  lethostigmus,  common  on  both  South  Atlantic 
and  Gulf  coasts ;  and  the  small  flounder  referred  to  above,  P.  albiguttus, 
which  also  occurs  on  both  Atlantic  and  Gulf  coasts.  For  the  market  no 
distinction  of  species  usually  is  made,  all  species  being  lumped  together 
as  flounders.  For  those  who  wish  to  tell  one  flounder  from  another  the 
following  suggestions  may  be  helpful.  In  southern  Florida  or  anywhere 
on  the  Gulf  coast  the  problem  is  simple,  for  only  two  species  occur — 
lethostigmus  and  albiguttus.  Lethostigmus  is  uniformly  dark  colored 
on  the  upper  surface,  while  albiguttus  has  a  number  of  conspicuous  spots, 
each  dark  with  a  white  ring  around  it.  From  North  Carolina  to  northern 
Florida  the  identification  is  a  little  more  difficult  because  three  species 


figure   20.-Tbe   southern   flounder,   found   from   Chesapeake   Bay   to   Texas,   is   the   common 
market  flounder  on  the  Gulf  coast. 


FISH    AND    SHELLFISH  39 

are  concerned.  However,  lethostigmus  is  still  easy  to  recognize  because 
it  lacks  spots.  As  between  dentatus  and  albiguttus,  both  of  which  bear 
the  ringed  spots,  remember  that  albiguttus  is  small  and  that  its  spots  are 
fewer — usually  three  placed  in  a  rough  triangle  plus  several  accessory 
spots,  compared  with  six  major  and  many  accessory  spots  in  dentatus. 
To  be  sure  of  his  identification  of  these  two  species,  however,  the  biologist 
counts  the  gill  rakers  and  fin  rays. 

All  flounders  have  flat,  compressed  bodies  and  a  peculiar  habit  of  lying 
on  their  sides.  They  are  not  born  with  this  habit  but  acquire  it  early  in 
life.  Several  amazing  structural  changes  accompany  the  transformation, 
most  important  of  which  is  the  fact  that  the  eye  on  the  under  side  moves 
across  the  forehead  and  comes  to  lie  beside  the  other  on  the  uppermost 
side  of  the  fish.  In  this  way  the  flounder  is  saved  the  inconvenience  of 
continuously  looking  into  the  mud  with  one  eye.  Some  flounders 
habitually  lie  on  the  left  side,  others  on  the  right.  The  southern  flounder, 
for  example,  lies  on  its  right  side.  Ordinarily,  the  under  side  becomes 
uniformly  light  in  color,  the  upper  side  uniformly  dark. 

Flounders  are  predatory  and  their  diet  consists  in  large  part  of  other 
fishes,  although  they  also  eat  shrimps,  crabs,  and  other  aquatic  animals. 
Often  they  lie  partly  buried  in  the  sand  with  only  the  eyes  exposed. 
Their  air  of  sluggishness  is  most  deceptive,  however,  for  when  a  fish 
passes  by  they  dart  up  with  surprising  rapidity  and  seize  it.  People  who 
have  kept  flounders  in  laboratory  tanks  say  that  they  will  strike  with 
great  force  at  a  finger  and  can  inflict  a  painful  wound. 

About  .Beaufort,  N.  C,  Hildebrand  and  Cable  (1930)  found  very 
young  flounders  off  Beaufort  Inlet  much  more  often  than  inside  the 
harbor,  from  which  they  inferred  that  most  or  all  of  the  spawning  takes 
place  at  sea.  The  principal  spawning  season  in  that  area  seems  to  be 
November  and  December.  The  young  soon  move  into  the  inside  waters, 
where,  like  the  adults,  they  live  on  or  near  the  bottom. 

Sharks  as  food  are  becoming  a  regularly  occurring  item  in  such  United 
States  markets  as  can  obtain  a  steady  supply.  Although  they  have  a 
long  and  honorable  history  of  utilization  in  other  countries,  sharks  had 
been  largely  neglected  by  the  American  public  as  food  until  wartime 
shortages  prompted  a  search  for  new  sources  of  protein  foods.  A  thriv- 
ing fishery  for  sharks  already  existed — especially  on  the  Pacific  coast 
?nd  in  North  Carolina  and  Florida  in  the  east — but  only  the  vitamin- 
rich  livers  and  the  hides  were  saved,  while  millions  of  pounds  of  meat 
were  being  discarded  at  sea.  The  first  experimental  shipments  of  shark 
meat  were  so  well  received  that  a  brisk  market  has  now  grown  up,  and 
the  supply  for  consumers  is  limited  only  by  the  ability  of  fishermen  to 
catch  and  process  the  sharks.  The  meat  is  placed  on  the  market  chiefly 
in  the  form  of  frozen  steaks  or  fillets.  The  present  heavy  demand  sug- 
gests that  shark  meat  will  continue  popular  after  the  war,  when  formerly 
plentiful  foods  have  been  restored  to  our  tables. 

The  chief  centers  of  shark  fishing  on  the  Atlantic  coast  are  Morehead 
City  in  North  Carolina  and  Mayport,  Salerno,  Cortez,  and  Key  West  in 
Florida.  Many  different  species  are  taken — probably  fifteen  to  twenty 
— with  the  leopard,  dusky,  brown,  sand,  hammerhead,  and  nurse  sharks 
being  among  the  most  important.  Gear  used  by  Florida  shark  fishermen 
resembles  the  long  trawl  lines  still  used  to  some  extent  in  New  England 
for  cod,  except  that  chains  are  substituted  for  the  lower  part  of  the  line 
because  of  the  sharp  teeth  and  the  strength  of  many  of  the  sharks. 

Sharks  are  classified  by  zoologists  as  fishes,  but  are  distinguished  from 
the  bony  fishes  by  the  fact  that  the  skeleton  is  not  ossified,  by  the 
presence  of  a  series  of  gill  slits  along  the  side  of  the  head,  and  by  other 


40 


CONSERVATION    BULLETIN    3  7 


less  conspicuous  anatomical  features.  The  general  habits  of  sharks  are 
very  similar  to  those  of  most  of  the  larger,  predatory  bony  fishes.  Their 
food  consists  largely  of  crabs,  lobsters,  small  fishes,  and  mollusks.  One 
of  the  largest  of  all  sharks,  the  basking  shark,  lives  on  the  minute 
surface  life  of  the  sea  which  it  strains  through  its  gills. 

Many  kinds  of  sharks  differ  from  most  bony  fishes  in  an  interesting 
respect — they  bring  forth  their  young  alive.  These  sharks  produce  eggs 
with  well  developed  shells  like  other  fishes,  the  difference  being  that  the 
eggs  hatch  within  the  body  of  the  mother,  instead  of  first  being  expelled. 
However,  while  the  cod,  for  example,  produces  millions  of  eggs,  the  shark 
may  give  "birth"  to  only  a  dozen  or  so  young.    This  great  difference  in 


Figure  21.— The  spiny  lobster  of  Florida,  also  called  the  sea  crayfish  or  rock  lobster,  is  a 
handsome  crustacean  mottled  with  blue,  rich  yellow,  and  brown  and  studded  with 
numerous   spines. 


FISH    AND    SHELLFISH  41 

fertility  is  probably  compensated  in  Nature's  scheme  by  a  higher  sur- 
vival rate.  The  young  sharks,  at  birth,  are  of  good  size,  active,  and 
obviously  ready  to  take  up  an  independent  existence. 

The  spiny  lobster  of  Florida,  Panulirus  argus,  is  not  closely  related  to 
the  New  England  lobster,  although  people  sometimes  confuse  the  two. 
One  of  the  chief  differences  is  the  fact  that  the  spiny  lobster  lacks  the 
giant  claws  that  are  a  characteristic  feature  of  the  New  England  lobster 
and  provide  a  considerable  quantity  of  choice  meat.  All  the  meat  of 
the  spiny  lobster  comes  from  the  flexible  abdomen  or  "tail."  The  frozen 
lobster  tails,  still  encased  in  their  shells  of  mottled  green  and  brown,  are 
the  common  market  form. 

Present  catches  of  the  spiny  lobster  amount  to  slightly  less  than  half 
a  million  pounds,  taken  chiefly  on  the  Florida  Reef  from  Miami  to  Dry 
Tortugas.  Although  this  species  occurs  as  far  north  as  Beaufort,  N.  C, 
United  States  fishermen  seldom  take  it  outside  the  State  of  Florida. 
The  best  fishing  grounds  are  along  the  southern  shores  of  the  reefs  and 
keys.  A  related  form  is  found  on  the  Pacific  coast,  south  of  Point 
Conception. 

The  average  market  size  of  the  Florida  spiny  lobster  is  about  9  to  10 
inches,  exclusive  of  the  long  antennae.  The  largest  specimen  of  which 
the  Fish  and  Wildlife  Service  has  authentic  record  measured  over  17 
inches  and  weighed  8  pounds. 

Most  of  the  catch  is  made  at  night  in  pots  which  the  lobsters  enter  as 
they  crawl  about  to  feed,  for  they  are  nocturnal  prowlers.  During  the 
day  they  lie  hidden  under  rock  ledges  or  among  sponges,  where  they  may 
be  detected  in  clear  water  by  the  protruding  antennae.  Coral  reefs, 
rocky  bottoms,  and  other  grounds  which  provide  good  cover  are  their 
favorite  haunts. 

The  hard-shell  clam,  Venus  mercenaria,  occurs  along  the  shores  of 
the  South  Atlantic  and  Gulf  coasts  all  the  way  from  North  Carolina  to 
Texas,  but  only  the  States  of  Florida  and  North  Carolina  have  important 
clam  fisheries.  Florida  produces  about  700,000'  pounds  and  North 
Carolina  500,000. 

The  southern  hard-shell  clam  is  the  same  species  as  the  well  known 
quahog  of  New  England  and  in  the  South  is  sometimes  called  the 
"round  clam"  or  "little  neck  clam."  On  the  Gulf  coast  of  Florida  the 
clams  grow  so  much  larger  than  along  the  South  Atlantic  coast  that 
they  are  considered  a  distinct  sub-species,  known  as  Venus  mercenaria 
mortoni.  The  shells  of  the  Florida  clams  may  be  5%  to  6  inches  long 
and  the  clam  may  weigh  several  pounds. 

Probably  the  largest  clam  bed  in  the  United  States  occurs  off  the  Gulf 
coast  of  Florida,  in  the  region  of  the  Ten  Thousand  Islands.  It  is  some 
40  miles  long  and  at  its  widest  parts  extends  out  into  the  Gulf  a  distance 
of  about  5  miles.  The  clams  are  distributed  over  about  150  square 
miles  of  bottom,  occurring,  as  might  be  expected,  more  thickly  in  some 
places  than  in  others. 

In  the  Florida  clam  fishery  two  kinds  of  gear  are  used,  both  of  which 
apparently  are  unique  in  the  United  States.  The  bulk  of  the  catch  is 
taken  by  a  dredge,  which  in  general  appearance  resembles  a  houseboat. 
The  digging  apparatus  picks  up  the  clams  on  its  curved  teeth  and  de- 
posits them  on  a  moving  wooden  escalator  or  conveyor  which,  in  turn, 
carries  the  clams  past  two  or  three  men  who  pick  out  the  live  clams. 
Dead  clams,  shells,  and  debris  are  automatically  discarded  when  the 
conveyor  passes  under  water  again.  The  dredge  is  at  one  end  of  a  1,200 
foot  cable  which  is  attached  to  an  800-pound  anchor.     It  moves  along 


42  CONSERVATION    BULLETIN    3  7 

slowly  by  taking  up  the  cable.  When  a  run  is  completed  the  cable  is 
released  and  the  dredge  drifts  back  with  the  wind  and  tide.  Each  run 
of  the  dredge  follows  a  course  like  the  radius  of  a  circle,  with  the  anchor 
at  the  center. 

Besides  the  dredge,  some  small  boats  are  operated.  Clam  diggers  in 
these  boats  use  a  long-handled  probe  which  bears  four  prongs  of  flexible 
steel.  The  probe  is  thrust  into  the  bottom  repeatedly  until  a  clam  is 
felt,  then  an  additional  thrust  secures  the  clam  between  the  prongs  so 
that  it  may  be  drawn  out. 

All  of  the  boats  now  working  the  clam  bed  are  operated  in  connection 
with  a  local  cannery,  where  the  entire  catch  is  processed.  In  North 
Carolina  another  clam  cannery  is  operated,  but  part  of  the  catch  in 
that  State  is  sold  fresh. 

Besides  the  hard  clam,  one  other  species — the  small  coquina  clam — ■ 
is  marketed  in  Florida  for  use  in  broths  and  chowders,  but  the  annual 
production  is  only  about  15,000  pounds. 

The  bay  scallop,  Pecten  irradians,  is  taken  commercially  in  only 
two  areas  on  the  South  Atlantic  and  Gulf  coasts — in  Carteret  County, 
N.  C,  and  on  the  Gulf  coast  of  Florida,  principally  about  Fort  Myers, 
Tampa,  and  Apalachicola. 

At  the  present  time,  the  Florida  production  is  much  the  larger, 
amounting  to  about  130,000  pounds,  compared  with  approximately  35,000 
in  North  Carolina.  The  small  quantity  of  scallops  now  taken  in  North 
Carolina  is  the  result  of  the  strange  disappearance,  about  ten  years  ago, 
of  a  marine  plant  known  as  eelgrass,  in  which  the  young  scallops  find 
necessary  protection.  When  the  eelgrass  was  killed  out  by  an  unex- 
plained disease  which  struck  suddenly  over  a  wide  area  along  the 
Atlantic  coast,  the  scallop  population  in  the  affected  areas  was  virtually 
wiped  out,  so  close  is  the  ecological  dependence  of  the  mollusk  on  the 
sheltering  vegetation.  The  blow  dealt  the  scallop  fishery  of  North  Caro- 
lina is  revealed  in  the  following  figures:  in  1927  the  catch  was  835,000 
pounds ;  in  1940,  34,000  pounds.  Eelgrass  does  not  grow  south  of  Bogue 
Sound  in  North  Carolina,  hence  the  Gulf  coast  fishery  was  not  affected. 
It  has,  however,  shown  some  decline  due  to  other  causes,  probably 
economic. 

Like  oysters  and  clams,  scallops  have  two  shells — hence  are  called 
"bivalves" — but  they  differ  considerably  in  their  habits  from  these 
sedentary  relatives.  Scallops  swim  about  actively  by  rapidly  opening 
and  closing  the  shells,  the  forcible  closing  expelling  a  jet  of  water  which 
furnishes  the  means  of  propulsion.  A  large  muscle  controls  the  shell 
movements,  and  this  muscle  (sometimes  called  the  "eye")  is  the  only 
part  of  the  scallop  that  is  eaten.  Europeans  eat  the  entire  scallop,  just 
as  whole  oysters  and  clams  are  eaten,  but  Americans  have  not  yet 
learned  to  eat  anything  but  the  excellently  flavored  muscle. 


FISH    AND    SHELLFISH  43 

APPENDIX 
The  Nutritive  Value  of  Fish  and  Shellfish 

Most  people  eat  fish  because  they  enjoy  their  varied  flavors  and  their 
adaptability  to  many  different  methods  of  preparation.  There  are 
further  benefits  from  eating  aquatic  food,  however,  for  fish  are  good 
natural  sources  of  calcium,  phosphorus,  iron,  and  copper,  and  provide 
protein  of  unexcelled  quality.  Some  species  also  furnish  vitamins  in 
appreciable  quantities  and  sea  fish  are  rich  in  iodine. 

Fish  are  an  important  source  of  proteins,  a  type  of  food  which  must 
be  included  in  the  diet  to  provide  the  elements  needed  to  grow  and 
repair  worn-out  body  tissues.  Some  proteins  are  complete  in  that  they 
supply  all  of  the  elements  needed;  others  are  incomplete  and  must  be 
supplemented  with  other  protein  foods  if  the  body  is  to  remain  in  normal 
health.  Fish  proteins,  like  those  in  beef,  pork,  and  other  meats,  are 
complete  in  themselves  and  proteins  of  this  type  should  supply  about 
one-third  of  the  daily  protein  requirement. 

Fish  are  an  excellent  source  of  most  of  the  minerals  which  the  body 
needs  to  develop  properly  and  perform  its  functions.  Calcium  and 
phosphorus  (without  which  proper  development  of  bones  and  teeth  is 
impossible)  occur  in  fish  fillets  in  about  the  same  quantities  as  in  beef 
round.  Marine  fishes  are  especially  rich  sources  of  iodine,  containing 
50  to  200  times  as  much  of  this  essential  element  as  any  other  food. 
Oysters,  shrimp,  and  crabmeat,  compared  with  milk,  provide  half  as 
much  calcium,  five  times  as  much  magnesium,  and  slightly  more  phos- 
phorus. Iron  and  copper,  which  build  up  the  hemoglobin  content  of 
the  blood  and  prevent  or  remedy  nutritional  anemia,  are  easily  obtained 
by  eating  most  fish.  Oysters  and  shrimp  are  the  best  known  sources  of 
these  two  minerals. 

Although  fish-liver  oils  have  long  been  recognized  as  first-class  sources 
of  vitamins  A  and  D,  it  is  less  widely  known  that  the  flesh  of  fish  is  also 
a  source  of  several  vitamins.  On  the  average,  daily  vitamin  require- 
ments could  be  obtained  from  ordinary  serving  portions  of  fish  to  the 
following  extent:  vitamin  A,  10  percent;  vitamin  D,  more  than  adequate 
amounts;  thiamin  (vitamin  Bi),  15  percent;  riboflavin  (vitamin  B2),  and 
nicotinic  acid  (another  element  of  the  vitamin  B  complex),  70  percent. 

General  Guides  for  Selecting  and  Preparing  Fish 

How  to  buy. — Insist  upon  freshness.  A  fresh  fish  may  be  recognized 
by  the  following:  firm  and  elastic  flesh,  scales  that  cling  to  the  skin  in 
most  species,  reddish  gills  free  from  disagreeable  odor,  eyes  bright  and 
full,  not  sunken.  In  selecting  shellfish  like  clams  and  oysters,  be  sure 
that  the  shells  are  tightly  shut,  indicating  that  the  animals  are  alive, 
unless  you  prefer  to  buy  the  meat  separately  as  shucked  shellfish.  Crabs 
and  lobsters  should  be  bought  alive  or  as  cooked  meat.  However,  un- 
cooked shrimp  may  be  bought  in  the  shell  provided  it  feels  firm  to  the 
touch.  Cooked  shrimp  is  sold  either  with  or  without  the  shell,  with  the 
heads  already  removed. 

When  to  buy. — In  general,  the  fish  of  any  species  are  of  highest  food 
quality  when  most  abundant,  for  at  these  periods  fishermen  are  making 
their  catches  in  the  shortest  time  and  shipping  them  promptly.  Usually, 
but  not  always,  fish  are  cheapest  when  most  abundant. 

Common  market  forms. — Fresh  (refrigerated)  fish  and  completely 
frozen  fish  should  be  equally  good  if  the  freezing  is  done  by  the  modern 


44  CONSERVATION    BULLETIN    3  7 

methods  now  well  known  to  the  industry.  Both  are  marketed  in  a 
variety  of  convenient  forms,  as  follows: 

Whole  or  round  fish  are  those  marketed  in  the  form  in  which  they 
come  from  the  water,  and  are  of  three  kinds:  fish  that  keep  as  well  or 
better  without  dressing,  small  fishes,  or  the  small  sizes  of  larger  species. 
Before  cooking,  whole  or  round  fish  are  eviscerated,  and  in  all  but  the 
very  small  sizes,  the  heads,  scales,  and  sometimes  the  fins  are  removed. 

Drawn  fish  are  those  marketed  with  only  the  entrails  removed.  To 
prepare  these  fish  for  cooking,  the  heads,  scales,  and  (if  desired)  the 
fins  are  removed,  and  the  fish  may  be  split  or  cut  into  serving  portions 
if  too  large  to  be  cooked  whole. 

Dressed  fish  have  had  the  head  and  entrails  removed  and  the  tail  and 
fins  may  be  cut  off.  If  dressed  fish  are  large,  they  may  be  cut  into  pieces 
in  preparation  for  cooking.  Very  large  dressed  fish  are  sometimes 
marketed  in  pieces. 

Steaks  are  slices  (usually  about  half  an  inch  thick)  cut  across  a  large 
dressed  fish. 

Fillets  are  meaty  slices  cut  lengthwise  from  the  sides  of  the  fish. 
Fillets  contain  no  bones  or  other  waste.  Their  weight  varies  with  the 
size  of  the  fish  from  which  they  are  cut. 

Stocks  are  crosswise  or  lengthwise  cuts  of  fillets. 

Canned  fish. — Besides  the  universally  familiar  canned  salmon,  tuna, 
and  sardines,  many  kinds  of  fish  are  canned  for  use  in  main  dishes, 
salads,  and  appetizers.  For  the  duration  of  the  war,  however,  the 
amount  of  canned  fish  available  for  civilians  will  be  considerably  less 
than  normal. 

Salt  or  Sm,oked  fish. — Tasty  variations  in  the  menu  are  provided  by 
salt  or  smoked  fish.  Salt  fish  ordinarily  requires  one-half  to  several 
hours'  soaking  before  further  preparation;  while  smoked  fish  usually  is 
ready  to  eat  as  it  is,  or  may  be  heated. 

Fat  content  of  fish. — For  best  results  in  preparing  a  fresh  fish,  it  is 
always  desirable  to  know  whether  it  is  fat  or  lean.  Fat  fish  are  especially 
suitable  for  baking,  and  may  also  be  broiled,  Avhile  lean  fish  are  best 
adapted  to  steaming,  boiling,  and  frying.  Medium-fat  fish  are  prepared 
like  the  lean,  or  may  be  dressed  with  strips  of  salt  pork  or  bacon  and 
baked.    Most  cook  books  classify  fish  as  follows: 

Fat  fish  are  those  containing  more  than  5  percent  fat.  Examples  are 
mullet,  Spanish  mackerel,  and  pompano. 

Lean  fish  are  those  containing  less  than  5  percent  fat.  Examples  are 
red  snapper,  sea  trout,  and  groupers. 

Sauces  and  garnishes. — The  attractiveness  of  almost  any  dish  con- 
sisting of  fish  will  be  increased  greatly  by  the  use  of  sauces  that  subtly 
enhance  or  complement  the  flavor.  Any  good  cook  book  contains  ex- 
cellent suggestions  as  to  the  choice  and  preparation  of  such  sauces.2 
Fresh  and  colorful  garnishes  also  do  much  to  create  a  dish  as  pleasing  to 
the  eye  as  to  the  palate,  thereby  whetting  the  appetite  and  helping  to 
make  the  serving  of  fish  a  pleasurable  and  often  repeated  experience. 


2  Sauces  for  seafoods.     Fishery  Leaflet   53.     Mimeographed,   4  pages,  may  be  obtained   on   request 
from  the  Fish  and  Wildlife  Service.  Chicago  54,  111 


FISH    AND    SHELLFISH  45 

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Conservation  Bulletin  No.  27.    24  pp.,  1943. 


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